Rhagovelia kawakamii hoberlandti Hungerford & Matsuda, 1961

Zettel, Herbert, Laciny, Alice & Freitag, Hendrik, 2020, Review of the genus Rhagovelia Mayr, 1865 (Insecta: Heteroptera: Veliidae) in the Palawan biogeographic region, the Philippines, Raffles Bulletin of Zoology 68, pp. 810-837 : 823

Rhagovelia kawakamii hoberlandti Hungerford & Matsuda, 1961

( Figs. 5A–D , 7G , 8D , 11E , 12B , 13C )

Rhagovelia (Rhagovelia) hoberlandti Hungerford & Matsuda, 1961: 260–261 . Type locality: Dimaniang (see notes under R. minutissima ), Busuanga (types not examined).

Rhagovelia kawakamii ( Matsumura, 1913) (partim) – Polhemus & Polhemus, 1988: 206 ( R. hoberlandti a synonym of R. kawakamii ).

Rhagovelia kawakamii forma hoberlandti – Zettel, 1996: 127–129.

Rhagovelia hoberlandti – Chen et al., 2005: 438.

Rhagovelia kawakamii hoberlandti – Freitag & Zettel, 2013: 64–65.

Philippine material reported by Zettel (1996). 3 apterous males, 4 apterous females from Busuanga, 13 road-km WNW Coron, Balulu Falls ; 6 apterous males, 9 macropterous males, 4 apterous females, 9 macropterous females from Busuanga , 5 km NW Coron, Mabintangen River ; 2 macropterous males, 1 apterous female from northern Palawan, Taytay , Binaluan ; 16 apterous males, 15 apterous females, 1 macropterous female from central Palawan, Cabayugan , 2 km N Sabang, 0–10 m a.s.l.; 1 apterous male, 1 apterous female from central Palawan, Cabayugan , stream in degraded forest near Sabang , 100 m a.s.l.; 8 apterous males, 3 macropterous males, 9 apterous females, 2 macropterous females from central Palawan, 9 km W Puerto Princesa, Iwahig , Balsahan River ; 2 apterous males, 3 apterous females from central Palawan, 20 km WSW Puerto Princesa, Montible River ; 6 apterous males, 6 apterous females from central Palawan, 20 km W Puerto Princesa, Tacduan Area , Tacduan River ; 1 apterous male, 4 macropterous males, 5 apterous females, 6 macropterous females from central Palawan, 2 km N Narra, Estrella Falls ; 3 apterous males, 2 apterous females from Balabac , Pasig; material in BMNH, BPBM, NHMW, UPLB, CZVA .

Figures

Fig. 5. Habitus of Rhagovelia kawakamii hoberlandti. A, apterous male; B, apterous female; C, macropterous male; D, macropterous female.

Fig. 7. Left parameres of Rhagovelia species, lateral view. A, R. minutissima; B, R. palawanensis; C, R. matillanoi, new species; D, R. estrella; E, R. abbreviata, new species; F, R. lansburyi; G, R. kawakamii hoberlandti; H, R. gapudi, new species.

Fig. 8. Proctiger of Rhagovelia species, dorsal view pilosity omitted. A, R. matillanoi, new species; B, R. abbreviata, new species; C, R. lansburyi; D, R. kawakamii hoberlandti; E, R. gapudi, new species.

Fig. 11. Area and distribution maps of Rhagovelia species. A, overview of the Palawan Region with major islands and towns, insert showing the location of the area within the Philippines as well as the northern part of Borneo belonging to Malaysia; B, allopatric distribution of the species of the R. sarawakensis group related with R. minutissima; C, allopatric distribution of the species of the R. sarawakensis group related with R. estrella; D, distribution of R. lansburyi; E, distribution of R. kawakamii hoberlandti; F, distribution of R. gapudi, new species.

Fig. 12. Habitats. A, a larger river at Napsan Road, kilometre 29, with high water level after heavy rain; R. palawanensis and R. estrella were recorded in quieter rock pools and between flooded vegetation. B, a small streamlet at Calatoboc Bridge over Northern National Highway was inhabited by R. kawakamii hoberlandti and R. gapudi, new species. Whereas the first species is euryoecious, R. gapudi, new species, prefers such small running waters.

Fig. 13. The Salakot Waterfalls and the stream below at Napsan Road, kilometre 43. In the various habitats, R. estrella, R. kawakamii hoberlandti, and R. lansburyi were recorded. Deep rock pools as shown in B are characteristic habitats for R. lansburyi.