Phanaeus (Notiophanaeus) achilles Boheman, 1858

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 206-207

Phanaeus (Notiophanaeus) achilles Boheman, 1858 Plate 45D

Phanaeus achilles Boheman, 1858: 42 (original description. Type locality: Insula Puna [= Isla Puna]).

Phanaeus achilles : Gemminger and Harold 1869: 1016 (catalog, distribution, written as Phanaeus Achiles Bohem); Nevinson 1892: 1 (catalog, distribution, written as Phanaeus Achiles Boheman); Gillet 1911a: 81 (catalog, distribution, written as Phanaeus Achiles Boh); d’ Olsoufieff 1924: 40 (characters in key); 97 (diagnosis), 151 (distribution); Blackwelder 1944: 209 (list of species from Latin America); Vulcano and Pereira 1967: 574 (characters in key); Ratcliffe et al. 2015: 197 (cited for Peru).

Phanaeus (Notiophanaeus) achilles : Edmonds 1994: 28 (characters in key), 30 (redescription), 99 (distribution); Edmonds and Zídek 2012: 10 (characters in key); Figueroa et al. 2014: 136 (distributional records from Peru); Chamorro et al. 2018: 97 (cited for Ecuador).

Phanaeus (Notiophanaeus) achilles achiles : Arnaud 2002a: 88 (diagnosis); Streit 2008: 8 (distribution, photography); Carvajal et al. 2011: 320-321 (cited for Ecuador); Krajcik 2012: 204 (complete list of species).

Phanaeus Achilles lydiae Arnaud, 2000: 8 (original description); Arnaud 2002a: 88 (diagnosis, distribution); Streit 2008: 9 (distribution, photography); Carvajal et al. 2011: 320-321 (cited for Ecuador); Krajcik 2012: 204 (complete list of species).

Phanaeus foveolatus Harold, 1880b: 152 (original description); Nevinson 1892: 4 (catalog, distribution); Gillet 1911a: 81 (cited as synonym of Phanaeus achilles Boheman, 1858); Blackwelder 1944: 209 (synonym of Phanaeus achilles Boheman, 1858); Edmonds 1994: 30 (cited as synonym of Phanaeus achilles Boheman, 1858); Arnaud 2002a: 88 (cited as synonym of Phanaeus achilles Boheman, 1858); Figueroa et al. 2014: 136 (cited as synonym of Phanaeus achilles Boheman, 1858).

Type specimens.

Phanaeus achilles Boheman, 1858. The holotype (♂) is deposited at the NHRS (see Edmonds 1994: 30). Locality: Puna, not examined.

Phanaeus foveolatus Harold, 1880. The holotype is deposited at the NMHU. Locality: Guayaquil, examined.

Holotype (♂): "Guayaquil. / Rusj [hw, green label]", "foveolatus / Harold [hw, green label]", "38678 [p]", "1405 [p]", "♂ HOLOTYPE [hw and p, red label]".

Phanaeus achilles lydiae Arnaud, 2000. The holotype (♂) is deposited at the CPFA (see Arnaud 2000: 8). Locality: Guayas, Los Ceibos Guayaquil (not examined). One paratype is deposited in MQCAZ, examined.

Paratype (♂): "ECUADOR (GUA) / Cerecita / 02 / 87 / P. Arnaud leg [p]", "Phanaeus achilles / lydiae / P. ARNAUD DET 96 / PARATYPE ♂ [p and hw, red margin]".

Distribution.

Ecuador and Peru.

Records examined.

EL ORO: 3 km S de Arenillas, 50 m (6 specimens CEMT); Arenillas, 15 m (75 specimens CEMT). GUAYAS: Cerecita (1 specimen MQCAZ); Guayaquil (1 specimen NMHU; 2 specimens CFPL); Los Ceibos Guayaquil (4 specimenes CEMT); Puna [= Isla Puna] (2 specimens NHRS). LOJA: Catamayo, Alamala, 1100 m (16 specimens MUTPL). MANABÍ: El Aromo, La Fabril, 290 m (1 specimen CEMT; 8 specimens MUTPL); El Aromo, Pueblo, 370 m (3 specimens MUTPL); Machalilla, 30 m (1 specimen CEMT); San Juan de Manta, 160 m (3 specimens MUTPL). UNDETERMINED PROVINCE: without specific locality (1 specimen NHRS).

Literature records.

GUAYAS: 40 km SW Guayaquil, 50 m ( Edmonds 1994: 99); 45 km W Guayaquil ( Edmonds 1994: 99); Bucay ( Edmonds 1994: 99); Env. Cerecita, Rte. Salinas ( Arnaud 2000: 8); Guayaquil ( Edmonds 1994: 99); Insula Puna [= Isla Puna] (Boheman 1858: 42; Edmonds 1994: 28); Los Ceibos, Guayaquil ( Arnaud 2000: 8; Arnaud 2002a: 88); Posorja, 0 m ( Arnaud 2000: 8; Edmonds 1994: 99). LOJA: Catamayo ( Edmonds 1994: 99); E35 9.5 km S. Catamayo 1211 m ( Streit 2008: 8); without specific locality ( Arnaud 2002a: 88); without specific locality ( Edmonds 1994: 99). MANABÍ: Montecristi ( Streit 2008: 9).

Temporal data.

Collected in January, February, March, April, May, June, October, November, and December.

Remarks.

Inhabits coastal lowland semi-deciduous forests from 5-370 m a.s.l. In the Andean region, it was recorded in the matorral dry montane forests from 1100-1210 m a.s.l. Collected manually and with pitfall traps baited with human feces.