Salvia cualensis J.G. González, 2012

González-Gallegos, Jesús Guadalupe & Castro-Castro, Arturo, 2012, Salvia cualensis and Salvia cualensis var. perezii (Lamiaceae), two new taxa from the Sierra de El Cuale, Jalisco, Mexico, Phytotaxa 74 (1), pp. 47-58 : 50-51

Salvia cualensis J.G. González , sp. nov. ( Figs. 1–2 , 4 )

Salvia jaimehintoniana primo adspectu maxima simile, sed foliis nunc anguste oblongo–lanceolatis nunc linearibus, inflorescentiis brevioribus; inflorescentiarum bracteis, calycibus, corollarum tubis, labiis et stylis longioribus differt.

Type:— MEXICO. Jalisco: Cabo Corrientes, 10.4 km por la brecha de El Tuito a Las Minas de Zimapán , a partir del km 4 al N de El Tuito, carretera a Puerto Vallarta , 1137 m, 20°21’27.14”N, 105°15’30.26”W, 12 August 2011 (fl, fr), J . G . González-G. 1078, A. Castro-Castro , R . Guerrero-H. & E. De Castro-Arce (holotype IBUG!, isotypes ENCB!, IEB!, MEXU!, XAL!, ZEA!) .

Perennial herb, erect, 20–70 cm tall, thickened roots, stems sparsely covered with retrorse and short glandularcapitate hairs towards the apex. Petioles 0–2.5 mm long, sparsely covered with retrorse hairs; leaf blades oblong-lanceolate to linear, (4.7–)8.6–14.4 × (0.3–) 0.9–1.7 cm, cuneate to rounded at the base, acute at the apex, the margin sparsely and shortly serrate to entire, both surfaces glabrous except the middle vein which is sparsely covered with short conical hairs and sometimes with tiny glandular capitate hairs, upper surface lustrous; usually with axillary fascicles of several leaves. Inflorescences (4–)8–17(–25) cm long, 4–7 verticillasters per inflorescence, verticillasters (2–)4- or 6(–8)-flowered, lowermost nodes 1.7–4.1 cm apart, the uppermost ones gradually crowded, floral axis covered with long eglandular and tiny glandular-capitate hairs (less than 0.5 mm long). Floral bracts ovate-lanceolate to lanceolate, (2–)14–18.6 × (1–) 4–6.7 mm, deciduous, truncate at the base, acute or long caudate at the apex, green and magenta tinged, the margin entire and bordered with erect hairs, the outer surface sparsely covered with appressed hairs, the inner one glabrous. Pedicel (2–) 2.7–4.4 mm long in flower, covered with long straight eglandular and tiny glandular-capitate hairs. Calyx (6–)7.9–11 × (3–) 4.4–6.5 mm, green, magenta or purple tinged at the upper lip, veins with long straight hairs and sometimes with tiny glandular-capitate hairs between them, internally covered with dark conical short hairs toward the apex, the upper lip 5-veined and entire, lobes of the lips acute, margin of the lips bordered with short simple hairs. Corolla white and the lips sky or dark blue, white nectar guides present on the lower lip, glabrous except the upper lip and the ventral surface of the lower one which are pilose; tube (6–)8.5–11 × 3.5–4.4 mm, straight or slightly constricted at the base, slightly ventricose, internally epapillate, upper lip (3.3–) 5.2–8 mm long, lower one (7–)11.1–15 × 6.8–11(15) mm. Stamens included; filaments 1.5–2.4 mm long; connectives 4.5–6.4(–7.5) mm long, ornate with an acute or rarely truncate short teeth at the insertion point with the filament; thecae 1.6–2.5 mm long; two staminodes present above and behind the insertion point of the filament to the corolla tube. Gynobasic horn 1.3–1.5 mm long; styles (9.5–) 12.4–13.6 mm long, pilose at the apex. Nutlets ovate, 2.8–3 × 2–2.4 mm, pale amber and irregularly dark brown reticulated, surface glabrous and smooth.

Figures

FIGURE 1. Distribution of Salvia cualensis and related taxa. Salvia cualensis var. cualensis (dots), S. cualensis var. perezii (triangles), S. heterotricha (squares), and S. jaimehintoniana (crosses).

FIGURE 2. Salvia cualensis var. cualensis. A) General aspect; B) Floral bract, outer surface; C) Flower; D) Corolla; E) Corolla section showing a filament and a staminode; F) Stamens; G) Apex of the style; H) Nutlets, dorsal and ventral view (based on J.G. González-G. 1078 et al.; drawn by J.G. González-G.).

FIGURE 4. Salvia cualensis var. perezii. A) Habit; C) Detail of flower. Salvia cualensis var. cualensis. B) Habit; D) Detail of flower. A–D by J.G. González-G.