Fagus gussonii MASSALONGO emend. KNOBLOCH et VELITZELOS

Teodoridis, Vasilis, Kvaček, Zlatko, Sami, Marco, Utescher, Torsten & Martinetto, Edoardo, 2015, Palaeoenvironmental Analysis Of The Messinian Macrofossil Floras Of Tossignano And Monte Tondo (Vena Del Gesso Basin, Romagna Apennines, Northern Italy), Acta Musei Nationalis Pragae Series B 71 (3 - 4), pp. 249-292 : 262

Fagus gussonii MASSALONGO emend. KNOBLOCH et VELITZELOS

Pl. 8, Fig. 9–12; Pl. 10, Fig. 4

Leaves elliptic to ovate, 47 to 110 mm long, 32 to 75 mm wide, base often asymmetric, rounded to widely cuneate with petiole up to 13 mm long, apex acute to attenuate, margin simply widely serrate, teeth regularly spaced, distinct, acute, venation simple craspedodromous, midrib straight, moderate, secondary veins straight, alternate, regularly spaced, tertiary veins straight to sinuous, alternate percurrent, venation of the higher orders regular polygonal reticulate, areolation well developed, 3- to 4-sided.

Such fossil foliage of beech has usually been assigned to Fagus gussonii based on the Neogene records from southern Europe ( Kvaček et al. 2002, Denk 2004).

M a t e r i a l. Leaf impressions (Monte Tondo MSF1608,

MSF1609, MSF1610, MSF1611, MSF1612, MSF1613,

MSF1614, MSF1615, MSF1616, MSF1617; Tossignano MSF25, MSF47, MSF258, MSF1532).