Orthocephalus saltator (Hahn)

Namyatova, Anna A. & Konstantinov, Fedor V., 2009, Revision of the genus Orthocephalus Fieber, 1858 (Hemiptera: Heteroptera: Miridae: Orthotylinae) 2316, Zootaxa 2316, pp. 1-118 : 72-74

Orthocephalus saltator (Hahn)

Figures 3, 8, 12, 16

Capsus saltator Hahn, 1835: 11 .

Capsus hirtus Curtis, 1838: 693 (junior primary homonym of Cimex hirtus Muller, 1776 ; syn. by Herrich–Schaeffer, 1853: 14).

Pachytoma major Costa, 1853: 56 (syn. by Puton, 1875: 38).

Orthocephalus saltator Fieber, 1861: 293 .

Globiceps infuscatus Garbiglietti, 1869: 190 (syn. by Puton, 1871: 425).

Orthocephalus ferrarii Reuter, 1891: 50 (syn. by Tamanini, 1977: 36).

Orthocephalus parvulus Reuter, 1891 NEW SYNONYMY.

DIAGNOSIS: Recognized by the dark brown hemelytra (fig. 3), yellowish tibiae often paler than femora, dark brown antennal segment I, aedeagus with two spicules, long spicule straight, equal in length to dorsal wall of theca, short spicule curved, half as long as long spicule (fig. 8), dorsal labiate plate with medial sclerites almost triangular, somewhat longer than sclerotized ring (fig. 16). Most easily confused with O. fulvipes and O. sefrensis , but differing from both in the straight rather than twisted long spicule of aedeagus (compare fig. 8 and figs. 7, 9), and from the former by the dark brown antennal segment I. Apparently not separable from these two species by the structures of the female genitalia (figs. 15, 16, 17). Also similar in appearance to O. bivittatus , but that species can be separated by having the long spicule shorter than the dorsal wall of the theca (fig. 7) and the medial sclerites distinctly longer than the sclerotized rings (fig. 14).

REDESCRIPTION: Male: Macropterous, length 4.2–6.4. COLORATION (fig. 3): Head: Dark brown to black, with black, sometimes pale brown, smooth spot near inner margin and with pale brown or yellowish stripe along posterior margin of eye, sometimes this color-pattern indistinct; antennal segment I dark brown to black, segment II dark brown, sometimes paler at base; labium dark brown to black. Thorax: Pronotum dark brown to black; evaporatorium at least partly yellowish or pale brown, paler than pleurites; scutellum dark brown to black; coxa dark brown to black, rarely forecoxa somewhat paler than middle and hind coxa; femur dark brown to black, rarely pale brown or yellowish apically; tibia yellowish or orange, apically and basally darkened; hemelytra dark brown to black; membrane uniformly pale brown. Abdomen: Uniformly dark brown to black. SURFACE AND VESTITURE: As in O. bivittatus (as in figs. 22 A-C, E, 23 A, D). STRUC- TURE AND MEASUREMENTS: Body 3.3–4.3× as long as width of pronotum. Head: Shape of head and eye as in O. bivittatus (as in figs. 20 B, 21 A, G), 1.2–1.3× as wide as high; vertex 1.6–2.3× as wide as eye; eye located a short distance from anterior pronotal margin; antennal fossa located close to inferior margin of eye; antennal segment I as long as or somewhat longer than diameter of eye, segment II 0.8–1.2× as long as basal width of pronotum, 1.1–1.6× as long as width of head; labium reaching middle coxa. Thorax: Pronotum 1.7– 2.3× as wide as long and 1.2–1.5× as wide as head, with posterior margin straight or slightly concave, lateral margin not carinate, calli slightly raised; hind femur relatively narrow, approximately 1.5× as wide as middle femur. GENITALIA: Aedeagus with two spicules, long spicule straight, not twisted, somewhat dilated in apical third, with gradually tapering apex, nearly equal in length to dorsal wall of theca; short spicule clawshaped, gradually curved, about 1.5× as short as long spicule ( Fig. 8); parameres as in fig. 12.

Female: Brachypterous, rarely macropterous; length 3.3–4.8. COLORATION: As in male. SURFACE AND VESTITURE: As in male. STRUCTURE AND MEASUREMENTS: As in male, body 2.5–3.5× as long as width of pronotum. Head: 1.1–1.2× as wide as high, vertex 1.7–2.1× as wide as eye; antennal segment II 0.8–1.2× as long as basal width of pronotum, 0.9–1.3× as long as width of head. Thorax: Pronotum 1.3–2.2× as wide as long and 0.9–1.3× as wide as head with posterior margin distinctly concave; length of hemelytra ranging in brachypterous specimens from reaching abdominal tergum VII to covering entire abdomen, usually covering abdominal tergum VIII, cuneus slightly delimited, claval commissure as long as inner claval margin, posterior and lateral margins of hemelytra rounded or posterior margin sometimes truncate. GENITALIA: Sclerotized rings oval, medial sclerites almost straight, twin-coned, with extreme bases twisted and variously shaped, truncate (as in O. fulvipes , fig. 15) or triangular apices, somewhat longer and narrower than sclerotized rings (fig. 16); posterior wall of bursa copulatrix with trapeziform sclerite uniformly sclerotized, incision on the anterior margin not reaching middle of sclerite, anterior angles of this sclerite somewhat dentate, posterior margin without sclerites (as in fig. 18, O. bivittatus ); vulva with curved sclerites forming a circle (as in fig. 19, O. vittipennis ).

DISTRIBUTION: Widely distributed in the western Palaearctic. The distributional range includes virtually all European countries and extends north to Norway, Sweden, Finland, Karelia Arkhangelsk, Perm, and Penza provinces of Russia, south to North Africa, Turkey, Caucasus, Iran, Kazakhstan, Altai, and Tuva, and east to Irkutsk province. This species was introduced to North America ( Henry and Kelton, 1986).

HOSTS: This species is undoubtedly polyphagous and has been collected from several plant families. It was recorded from the following species: Asteraceae : Chrysanthemum sp. ( Reuter, 1891) , Centaurea maculosa Lam. and Cichorium intybus L. ( Henry and Kelton, 1986), Centaurea sp. and Hieracium pilosella L. ( Wagner, 1974), Hieracium alpinum L., H. villosum Jacq. , H. racemusum Waldst. et Kit. ( Tamanini, 1981, 1982), Erigeron and Crepis spp. ( Tamanini, 1981) ; Fabaceae : Vicia sp. and Trifolium sp. ( Reuter, 1891) , Ononis sp. ( Saunders, 1892) , Spartium , including Spartium junceum L. ( Tamanini, 1981), Lotus sp. ( Seidenstücker, 1959) ; Lamiaceae : Thymus serpyllum L. ( Reuter, 1891). Ehanno (1960) collected O. saltator from species of Poaceae and Linnavuori (1999) from unspecified species of Asteraceae . Some specimens from ZISP were collected from Spiraea sp. (Rosaceae) by I. M. Kerzhner and Salix sp. (Salicaceae) by an unknown collector.

DISCUSSION: In this species, sclerites adjacent to the sclerotized rings vary in shape and can be very similar to those of O. fulvipes (see fig. 15). Females of O. saltator are difficult to separate from O. fulvipes and O. sefrensis as both external characters and genitalic structures are similar. Refer to O. sefrensis for additional discussion of the issue.

According to the original description ( Reuter, 1891), O. parvulus is related to O. saltator , but differs in the smaller size, thinner antennae, antennal segments III and IV combined longer than segment II, less convex eyes, and more inclined cuneal fracture. Wagner (1974) also suggested that these two species can be distinguished by the body length and ratio of vertex to eye. The careful examination of all available material has showed that all mentioned characters vary greatly and are not useful in the separation of O. saltator and O. parvulus . No distinctions were found in the structure of both male and female genitalia. Therefore, O. parvulus is synonymized with O. saltator .

Two syntypes of O. ferrarii and a single syntype of O. parvulus , belonging to the Reuter collection, Finnish Museum of Natural History, were studied in the course of this study. Here we designate the lectotypes of these two species. The lectotype of O. ferrarii , bears the locality label “Stazzano,” date and collector unknown and the type label “Spec. typ No 3504 Orthocephalus ferrari Reut. ” The lectotype of O. parvulus has the locality label “ Smyrna, 3.V.1871, Krueper” and the type label “Spec. typ No 3510 Orthocephalus parvulus Reut. ” The scientific names of species on both type labels are written in the Reuter’s hand.

SPECIMENS EXAMINED: Lectotype of Orthocephalus ferrarii : ITALY: Piedmont: Stazzano , 44.72758 ° N 8.87251 ° E, 231 m, Jun 1881, collector unknown, 1♂ ( AMNH _ PBI 00250631 ) ( MZH) . Lectotype of Orthocephalus parvulus : TURKEY: Izmir: Izmir [Smyrna], [Mont Jamanl.], 38.431 ° N 27.146 ° E, 03 May 1871, Krueper , 1♂ ( AMNH _ PBI 00268076 ) ( MZH) . Paralectoype of Orthocephalus ferrarii : ITALY: Piedmont: Stazzano , 44.72758 ° N 8.87251 ° E, 231 m, Jun 1881, collector unknown, 1♂ ( AMNH _ PBI 00258514 ) ( MZH) .

ADDITIONAL MATERIAL: ARMENIA: 26.6 km [25 verst] N KnyskalaTur, 28 Jul 1916, Andrievskiy, 2♀ (AMNH_PBI 00261477, AMNH_PBI 00261478) (ZISP). Allaverdi, 41.11666 ° N 44.85 ° E, 15 Aug 1930, Korinek, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261275) (ZISP); 03 Sep 1930, Korinek, ( Chloanthaceae ), 1♂ (AMNH_PBI 00260672) (ZISP). Ankavan [Mis-Chana], 40.61666 ° N 44.46666 ° E, 12 Aug 1932, Korinek, ( Chloanthaceae ), 2♂ (AMNH_PBI 00261048, AMNH_PBI 00261049) (ZISP); 14 Aug 1932, Korinek, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261274) (ZISP). Arich [Kipchag] NW Slops of Aragatz [Alagez] Mt., 40.61666 ° N 43.98333 ° E, 28 Jul 1935, W. Richter, 1♀ (AMNH_PBI 00261480) (ZISP). Between Meghri and Legvaz [Lishkvaz], 38.91666 ° N 46.21666 ° E, 20 Jul 1932, Korinek, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261273) (ZISP). Dilijan [Delizhan], 40.73333 ° N 44.86666 ° E, Melinyants, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261331) (ZISP); 22 Jul 1935, Ter-Minasyan & Richter, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261332) (ZISP); 24 Jul 1935, Ter-Minasyan & Richter, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261333) (ZISP). Elenovka nr Gokcha, Yerevan Dist., 40.55 ° N 44.95 ° E, 25 Jul 1927, Dyakonov, 1♂ (AMNH_PBI 00261354) (ZISP). Ellidzha, Kotayk' distr., 40.275 ° N 44.66056 ° E, 13 Jul 1954, Ter-Minasyan, 1♀ (AMNH_PBI 00261265), 1 juvenille (AMNH_PBI 00261034) (ZISP). Goris, Geruschay River, Elizavetpol Government, 39.5 ° N 46.33333 ° E, 08 Jul 1913, Bartenev, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261495) (ZISP). Karmrakar, 40.86666 ° N 43.88333 ° E, 12 Jun 1955, Loginova, 1♂ (AMNH_PBI 00259359) (ZISP); 14 Jun 1955, Loginova, 1♀ (AMNH_PBI 00261491) (ZISP); 16 Jun 1955, Loginova, 2♀ (AMNH_PBI 00261489, AMNH_PBI 00261490) (ZISP). Kushchibelyak, 110 km SE of Erevan, 39.799 ° N 45.6625 ° E, 17 Jul 1934, Ter-Minasyan, 1♂ (AMNH_PBI 00261033), 2♀ (AMNH_PBI 00261259, AMNH_PBI 00261260) (ZISP); 19 Jul 1934, Ter-Minasyan, 6♀ (AMNH_PBI 00261256 through AMNH_PBI 00261258, AMNH_PBI 00261261 through AMNH_PBI 00261263) (ZISP). Semenovka, Kamo [Novobayazet] distr., 40.651 ° N 44.899 ° E, 26 Jul 1954, Ter-Minasyan, 1♀ (AMNH_PBI 00261264) (ZISP). Yerevan, 40.2 ° N 44.53333 ° E, 20 Jul 1932, Korinek, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261276) (ZISP). AZERBAIJAN: Nakhichevan Prov.: Tyllyak on Gilyanchay River, 38.91 ° N 45.82 ° E, 01 Aug 1933, Znoiko, ( Chloanthaceae ), 3♀ (AMNH_PBI 00261334 through AMNH_PBI 00261336) (ZISP). Arpa-chay River, 39.4675 ° N 44.93444 ° E, 02 Jul 1937 - 06 Jul 1937, Ryabov, 1♀ (AMNH_PBI 00261488) (ZISP). Zarjal, Gansa [Elizavetpol], Tranccaucasia Gub., 40.68333 ° N 46.33333 ° E, 14 Aug 1913, Arzimowicz, 1♀ (AMNH_PBI 00268213) ( Chloanthaceae ), 1♀ (AMNH_PBI 00261485) (ZISP). BELARUS: Ambrosovichi, 55.2 ° N 29.666 ° E, 20 Jul 1924, Birulya, 1♂ (AMNH_PBI 00261201) (ZISP); 16 Jun 1926, Birulya, 1♂ (AMNH_PBI 00261202) (ZISP). Korolevo nr Vitebsk, 55.13333 ° N 30.5 ° E, 09 Jul 1905, Birulya, 1♀ (AMNH_PBI 00261319) (ZISP). Lavy, 10 verst S Sirotino, 55.25 ° N 29.61666 ° E, 09 Aug 1924, Birulya, 1♀ (AMNH_PBI 00261318) (ZISP). Zamostochye St., Vitebsk distr [former Senno uezd], 55 ° N 30.15 ° E, 12 Jun 1905, Birulya, 8♂ (AMNH_PBI 00261114 through AMNH_PBI 00261121), 4♀ (AMNH_PBI 00261110 through AMNH_PBI 00261113) (ZISP). BOSNIA HERCEGOVINA: Igman Mts., nr Saraevo, Bosnia, 43.76666 ° N 18.25 ° E, 30 Jul 1929, Martino, 1♂ (AMNH_PBI 00260684) (ZISP). BULGARIA: Dragalevtsi, Sofia, 42.61666 ° N 23.3 ° E, 09 Jul 1953, Martino, 3♀ (AMNH_PBI 00261266 through AMNH_PBI 00261268) (ZISP). Sandanski, 41.558 ° N 23.288 ° E, 29 May 1965, M. Josifov, 2♂ (AMNH_PBI 00263428, AMNH_PBI 00268067), 3♀ (AMNH_PBI 00240602, AMNH_PBI 00268068 through AMNH_PBI 00268069) (MHNG). ESTONIA: Sillamae [Sillamagi], 59.405 ° N 27.766 ° E, 24 Jun 1900, Bianchi, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261345) (ZISP); 28 Jun 1900, Bianchi, ( Chloanthaceae ), 1♂ (AMNH_PBI 00152909) (ZISP). GEORGIA: Abul-Sansarskiy Mts. Ridge, 41.51 ° N 43.66 ° E, 06 Aug 1931, Dyakonov, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261338) (ZISP). Aleksandrsgilf on the Khrami River, 41.59 ° N 44.08 ° E, 1569 m, 15 Aug 1931, Dyakonov, ( Chloanthaceae ), 2♀ (AMNH_PBI 00261339, AMNH_PBI 00261340) (ZISP). Benara, 19 km W Akhaltsykhe, 41.65 ° N 42.815 ° E, 12 Jun 1949, A. N. Kiritshenko, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261341) (ZISP); 20 Jun 1949, A. N. Kiritshenko, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261342) (ZISP). Gometsari canyon [Gometrits-Kheoba], valley of Aragvi River, 42.30306 ° N 44.81333 ° E, 11 Jul 1949, A. N. Kiritshenko, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261343) (ZISP). Lentekhi, Svanetia, 42.78333 ° N 42.7 ° E, 08 Aug 1957, Akramovskaya, 1♂ (AMNH_PBI 00260680) ( Chloanthaceae ), 1♂ (AMNH_PBI 00261680) (ZISP). Sasashi, Mestiyskiy Dist., Svanetiya, 42.76666 ° N 42.96666 ° E, 03 Aug 1957, Akramovskaya, 1♀ (AMNH_PBI 00235297) ( Chloanthaceae ), 4♂ (AMNH_PBI 00261676 through AMNH_PBI 00261679), 1♀ (AMNH_PBI 00261059) (ZISP). GERMANY: Bayern: Eichstaett Umgeb., 07 Jul 1959, Seidenstücker, 1♀ (AMNH_PBI 00230886) (ZISP); 04 Jul 1966, Seidenstücker, 2♂ (AMNH_PBI 00151515, AMNH_PBI 00268034) (ZISP). Mecklenburg-Vorpommern: Stettin Lake, 53.81 ° N 13.86 ° E, 20 Jun 1908, Dyakonov, 1♂ (AMNH_PBI 00260685), 1♀ (AMNH_PBI 00261066) (ZISP). GREECE: (Eastern) Crete: Lasithion: Koutsounara, 35 ° N 25.81666 ° E, 29 Apr 1999, A. Matocq, 1♂ (AMNH_PBI 00264058), 2♀ (AMNH_PBI 00264063) (MATOCQ). Psiloritis Mt., 35.25 ° N 24.75 ° E, 1420 m, 01 May 1999, A. Matocq, 1♂ (AMNH_PBI 00263373) (MATOCQ). Central Greece: Khalkis [Chalkis], Euboea, 38.46 ° N 23.6 ° E, Apr 1926, Holtz, 1♂ (AMNH_PBI 00261369) (ZISP). Dodecanese Islands: Olympos, 35.73333 ° N 27.18333 ° E, 03 Jun 1998, A. Carapezza, 1♂ (AMNH_PBI 00237601), 1♀ (AMNH_PBI 00268075) (AC). Makedonia: 10 km S Kalokastron, Mecedonien, 40.92 ° N 23.34 ° E, 03 Jun 1972, Eckerlein, 4♀ (AMNH_PBI 00268059 through AMNH_PBI 00268062) (MHNG). Kavala, 40.94 ° N 24.42 ° E, 04 May 1974, Eckerlein, 1♂ (AMNH_PBI 00226421), 1♀ (AMNH_PBI 00268063) (MHNG). Peloponnese: Achaea (Achaia) Co. : Erimanthos Mt, Minas, 37.97 ° N 21.8 ° E, 1200 m, 17 Jun 1990, R. Linnavuori, 1♂ (AMNH_PBI 00268094) (LCRT). Arkadia Co.: Between Levidhion [Lavidi] and Kandhila [Kandila], 37.73 ° N 22.34 ° E, 13 Jun 1989, R. Linnavuori, 1♀ (AMNH_PBI 00268093) (LCRT). Killini Mt. nr Ayios Nikolaos, 37.92 ° N 22.43 ° E, 12 Jun 1989, R. Linnavuori, 1♂ (AMNH_PBI 00268092) (LCRT). Laconia (Lakonias) Co. : Kiparission [Kyparissia], 36.96 ° N 22.98 ° E, 10 Apr 1956 - 15 Apr 1956, Schubert, 1♀ (AMNH_PBI 00268070) (MHNG); 02 May 1956, Schubert, 3♂ (AMNH_PBI 00152914, AMNH_PBI 00268064, AMNH_PBI 00268066), 1♀ (AMNH_PBI 00268065) (MHNG). Messinia (Messenia, Messinias) Co.: Manesis [Manesi], 37.08 ° N 21.9 ° E, 17 Jun 1990, R. Linnavuori, 1♀ (AMNH_PBI 00268095) (LCRT). Thessalonikis: approx. 50 km east of Thessaloniki, 38.62482 ° N 24.11563 ° E, 30 Jun 1974, Eckerlein, 1♂ (AMNH_PBI 00268058), 2♀ (AMNH_PBI 00268039, AMNH_PBI 00268057) (MHNG). Arcadie, Vitina, 37.66666 ° N 22.16666 ° E, 06 May 1994, Ph. Magnien, J. Pericart & A. Matocq, 2♂ (AMNH_PBI 00252352), 2♀ (AMNH_PBI 00248985) (MATOCQ). Argolie, Paralia, Astros, 37.4 ° N 22.71666 ° E, 05 May 1994, Ph. Magnien, J. Pericart & A. Matocq, 1♂ (AMNH_PBI 00248984), 1♀ (AMNH_PBI 00248984) (MATOCQ). Corinthe, Loutrakion [Loutraki], 37.96666 ° N 22.96666 ° E, 29 Apr 1994, A. Matocq and J. Pericart, 1♀ (AMNH_PBI 00263376) (MATOCQ). Corinthe, Perakhora, 38.03333 ° N 22.95 ° E, 24 Apr 1994, Ph. Magnien, J. Pericart, E. Heiss and A. Matocq, 2♂ (AMNH_PBI 00262906) (MATOCQ). Fokida, Ilia [Ilea], 38.85 ° N 23.11 ° E, 08 Jun 1996, S. Doguet, Ph. Ponel and A. Matocq, 1♀ (AMNH_PBI 00264062) (MATOCQ). Laconie, Elos Plage, 36.81666 ° N 22.68333 ° E, 30 Apr 1994, Ph. Magnien, J. Pericart, E. Heiss and A. Matocq, 2♂ (AMNH_PBI 00262907) (MATOCQ). Laconie, Khania [Hania], 36.88333 ° N 22.51666 ° E, 02 May 1994, Ph. Magnien, J. Pericart & A. Matocq, 2♂ (AMNH_PBI 00263368, AMNH_PBI 00263369), 3♀ (AMNH_PBI 00262908, AMNH_PBI 00264060) (MATOCQ). Laconie, Magne, Cape Matapan [Tenaro], 36.63333 ° N 22.8 ° E, 02 May 1994, Ph. Magnien, J. Pericart & A. Matocq, 1♂ (AMNH_PBI 00263370) (MATOCQ). Laconie, Monemvasia, 36.68333 ° N 23.03333 ° E, 04 May 1994, Ph. Magnien, J. Pericart & A. Matocq, 1♂ (AMNH_PBI 00263426), 1♀ (AMNH_PBI 00263426) (MATOCQ); 17 Apr 1997, J.M. Maldes and A. Matocq, 3♂ (AMNH_PBI 00264061), 1♀ (AMNH_PBI 00252544) (MATOCQ). Peloponese, Agiorgitika, 37.48333 ° N 22.46666 ° E, 23 May 1995, Doguet, 1♂ (AMNH_PBI 00263372) (MATOCQ). IRAN: Ardebil: Khemes, 37.52 ° N 48.63 ° E, 07 Jul 2002 - 09 Jul 2002, R. Linnavuori, 1♂ (AMNH_PBI 00268091) (LCRT). Gilan: 20-30 km E Khalkhal, E. Azerbajan, 37.62 ° N 48.8 ° E, 01 Jul 1996 - 21 Jul 1996, R. Linnavuori, 2♂ (AMNH_PBI 00268084, AMNH_PBI 00268085) (LCRT). Between Deylaman and Barasar, 36.82 ° N 49.82 ° E, 23 Jul 1996, R. Linnavuori, 1♀ (AMNH_PBI 00268086) (LCRT). Khuzestan: Beuran nr Seresht, 32.33 ° N 49.69 ° E, 09 Jul 1914, Nesterov, 1♂ (AMNH_PBI 00261364) (ZISP). ITALY: Lombardy: Bormio, 46.46 ° N 10.36 ° E, 1933, E. Passauro, 1♀ (AMNH_PBI 00261512) (ZISP). Milano, 45.46666 ° N 9.16666 ° E, 27 Jun 1947, Rubtsov, 1♂ (AMNH_PBI 00260682) (ZISP). Varese, 45.82 ° N 8.83 ° E, Jun 1934, Holtz, 2♂ (AMNH_PBI 00261367) (ZISP). Roma: Rome, 41.86666 ° N 12.48333 ° E, 30 May 1947 - 31 May 1947, Rubtsov, 1♂ (AMNH_PBI 00260683), 2♀ (AMNH_PBI 00261064, AMNH_PBI 00261065) (ZISP). Sicily: San Martino de ile Scale (Palermo), 38.2 ° N 15.38 ° E, 18 May 1961, Eckerlein, 1♀ (AMNH_PBI 00267510) (MHNG). KAZAKH- STAN: Kostanay Prov.: 250 km S Kustanai, Ak-Suat Lake, 51.41 ° N 64.47 ° E, 25 Jun 1946, Formozov, 1♀ (AMNH_PBI 00261508) (ZISP). North Kazakhstan Prov.: Between Shchuchinsk [Shchuch'ye] and Barmashino [Barmashi], 52.93333 ° N 70.21666 ° E, 05 Jul 1928, Filip'ev, 8♂ (AMNH_PBI 00261020 through AMNH_PBI 00261027), 3♀ (AMNH_PBI 00261246 through AMNH_PBI 00261248) (ZISP). Zhambul Prov.: Achi-Say, Teresakan river, Karatau Mts Ridge, 43.55417 ° N 68.88889 ° E, 01 Jun 1936, A.K. Lukyanovich, 1♀ (AMNH_PBI 00261505) (ZISP). LATVIA: Nider-Bartau, Grobina Uezd, Kurlyandiya, 56.54 ° N 21.16 ° E, Jul 1896, Beier, 1♀ (AMNH_PBI 00261503) (ZISP). Stalzen nr Vindava, Kurlyandskaya Government, 57.44111 ° N 21.60833 ° E, 16 Jul 1911, Birulya, 1♀ (AMNH_PBI 00261507) (ZISP). NETHER- LANDS: Drente: Schoonlo, 52.91 ° N 6.7 ° E, 16 Jul 1981, Aartsen, 1♂ (AMNH_PBI 00268031) (ZMAN). Limburg: Broekhuizen, 51.49 ° N 6.16 ° E, 03 Jul 1942, Gravestein, 1♂ (AMNH_PBI 00268028) (ZMAN). Vijlen, 50.79 ° N 5.96 ° E, 21 Jul 1965, Woudestra, 1♂ (AMNH_PBI 00157556), 1♀ (AMNH_PBI 00268030) (ZMAN). POLAND: Bialystok [Belostok], 53.128 ° N 23.151 ° E, 02 Jul 1913, Trzheskovskiy, 1♀ (AMNH_PBI 00261469) (ZISP). Bodzentyn, 50.93333 ° N 20.95 ° E, 20 Jul 1895, Jakobson, 1♀ (AMNH_PBI 00261282) (ZISP). Khabirov, Kalish Dist., 51.75 ° N 18.08333 ° E, 07 Jul 1907, Jachevski, 2♀ (AMNH_PBI 00261060, AMNH_PBI 00261061) (ZISP); 09 Jul 1907, Jachevski, 1♂ (AMNH_PBI 00260681) (ZISP); 18 Jul 1907, Jachevski, 1♀ (AMNH_PBI 00261062) (ZISP); 31 Jul 1908, Yachevskiy, 1♀ (AMNH_PBI 00261063) (ZISP). ROMANIA: Slopes of Pietros Mt., Zakarpatskaya Prov., 47.83 ° N 24.61 ° E, 21 Jul 1953, Roshko, 4♂ (AMNH_PBI 00261324 through AMNH_PBI 00261327) (ZISP). RUSSIAN FEDERATION: Altai Rep.: 20 km SE of Ongudai, 50.627 ° N 86.34 ° E, 17 Jul 1964, I. M. Kerzhner, Spiraea sp. (Rosaceae) , 1♂ (AMNH_PBI 00261362) (ZISP). Kebezen' nr Teletskoe Lake, 51.91 ° N 87.12 ° E, 18 Jun 1912, Yurganova, 1♂ (AMNH_PBI 00261361) (ZISP); 26 Jun 1912, Yurganova, 1♂ (AMNH_PBI 00261360) (ZISP). Kumurtuk, Chulyshman River Valley, 51.33333 ° N 87.73333 ° E, 07 Jul 1909, Emeljanov, 1♀ (AMNH_PBI 00261501) (ZISP). Verkhny Uymon - Kotanda, Valley of Katun' River, Biysk Uezd, 50.21666 ° N 85.71666 ° E, 10 Jul 1897 - 18 Jul 1897, Silantev, 1♂ (AMNH_PBI 00261368) (ZISP). Arkhangelsk Prov.: Left bank of Severnaya Dvina River, oppos. Kotlas, 61.25 ° N 46.51666 ° E, 27 Jul 1942, Stark, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261222) (ZISP); 29 Jul 1942, Stark, ( Chloanthaceae ), 2♂ (AMNH_PBI 00261009, AMNH_PBI 00261010), 5♀ (AMNH_PBI 00261217 through AMNH_PBI 00261221) (ZISP); 02 Aug 1942, Stark, ( Chloanthaceae ), 4♀ (AMNH_PBI 00261208 through AMNH_PBI 00261211) (ZISP); 04 Aug 1942, Stark, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261008), 4♀ (AMNH_PBI 00261212 through AMNH_PBI 00261215) (ZISP); 14 Aug 1942, Stark, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261216) (ZISP). Right bank of Severnaya Dvina River, 11 km E of Kotlas, 61.25 ° N 46.85 ° E, 18 Jul 1942 - 19 Jul 1942, Stark, ( Chloanthaceae ), 2♀ (AMNH_PBI 00261223, AMNH_PBI 00261224) (ZISP). Shipitsyno, nr Kotlas, 61.285 ° N 46.509 ° E, 13 Jul 1942, Stark, 1♂ (AMNH_PBI 00261296) (ZISP); 16 Jul 1942, Stark, 1♀ (AMNH_PBI 00260933) (ZISP). Bashkortostan Rep.: Aksenovo St., 53.91666 ° N 54.56666 ° E, 14 Jul 1908 - 16 Jul 1908, A. Grigorev, 1♀ (AMNH_PBI 00261291) (ZISP). Bryansk Prov.: Bryansk, 53.25 ° N 34.4 ° E, 06 Jul 1927, Stark, 1♂ (AMNH_PBI 00261189) (ZISP); 14 Jul 1927, Stark, 1♂ (AMNH_PBI 00261188), 1♀ (AMNH_PBI 00261191) (ZISP). Petrovskoe, [former Mtsenskiy uyezd of Orel government], 52.78333 ° N 33.08333 ° E, 06 Jun 1897, Favorskiy, 4♀ (AMNH_PBI 00261190, AMNH_PBI 00261352, AMNH_PBI 00261466, AMNH_PBI 00261509) (ZISP). Chelyabinsk Prov.: Between Alekseevskiy [Alekseevskaya] and Mikhaylovka, Troitskiy Uezd., 53.15 ° N 59.76666 ° E, 08 Jun 1916, N. Kuznetsov, 1♂ (AMNH_PBI 00261366) (ZISP). Il'menskiy Nature Reserve, 54.99 ° N 66.09 ° E, 13 Jul 1959, V.I. Tobias, 1♂ (AMNH_PBI 00261359) (ZISP). Chuvashia Rep.: Tsivilsk, 55.85 ° N 47.46666 ° E, 14 Jul 1875, Cherkanovskiy, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261337) (ZISP). Dagestan Rep.: Buyanaksk [Temir-Khan-Shura], 42.81666 ° N 47.11666 ° E, 20 Jul 1944, Ryabov, 1♀ (AMNH_PBI 00261487) (ZISP). Derbent, 42.05 ° N 48.28333 ° E, 11 Jun 1904, K. Satunin, 1♀ (AMNH_PBI 00261510) (ZISP); 07 Sep 1931, Ryabov, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261486) (ZISP). Khunzakh, 42.55 ° N 46.75 ° E, 22 Jul 1924, Ryabov, 2♀ (AMNH_PBI 00260902, AMNH_PBI 00260903), 2 juvenille (AMNH_PBI 00260993, AMNH_PBI 00260994) (ZISP); 23 Jul 1924, Ryabov, 7♂ (AMNH_PBI 00156269, AMNH_PBI 00260988 through AMNH_PBI 00260992, AMNH_PBI 00264294), 11♀ (AMNH_PBI 00227955, AMNH_PBI 00260892 through AMNH_PBI 00260901) (ZISP). Ingushetia Rep.: Between Salgi and Egochkal, 42.78 ° N 44.68 ° E, 25 Jul 1927, A. N. Kiritshenko, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261328) (ZISP). Between Salgi and Pass to Valley of Assa River, 42.78 ° N 44.68 ° E, 30 Jul 1927, A. N. Kiritshenko, ( Chloanthaceae ), 2♂ (AMNH_PBI 00261356, AMNH_PBI 00261357), 2♀ (AMNH_PBI 00261483, AMNH_PBI 00261484) (ZISP). Salgi, 42.78333 ° N 44.85 ° E, 1665 m, 22 Jul 1927, A. N. Kiritshenko, 1♂ (AMNH_PBI 00261092) (ZISP); 27 Jul 1927, A. N. Kiritshenko, 2♂ (AMNH_PBI 00261089, AMNH_PBI 00261090), 1♀ (AMNH_PBI 00261094) (ZISP); 29 Jul 1927, A. N. Kiritshenko, 2♂ (AMNH_PBI 00261090, AMNH_PBI 00261091) (ZISP); 31 Jul 1927, A. N. Kiritshenko, 1♂ (AMNH_PBI 00239070) (ZISP); 03 Aug 1927, A. N. Kiritshenko, 2♀ (AMNH_PBI 00261093, AMNH_PBI 00261095) (ZISP). Irkutsk Prov.: Bunbuy, 56.38306 ° N 99.02861 ° E, 22 Jun 1915 - 24 Jun 1915, Varaksina, ( Chloanthaceae ), 4♂ (AMNH_PBI 00261011 through AMNH_PBI 00261014), 8♀ (AMNH_PBI 00261225 through AMNH_PBI 00261232) (ZISP); 25 Jun 1915 - 28 Jun 1915, Varaksina, ( Chloanthaceae ), 2♀ (AMNH_PBI 00261233, AMNH_PBI 00261234) (ZISP). Khayryuzovka, Balaganskaya steppe, 53.76666 ° N 103.35 ° E, 17 Jul 1932, Zakhvatkin, 1♀ (AMNH_PBI 00261499) (ZISP). Mol'ka (eastern), Balaganskaya steppe, 54.5 ° N 103.26666 ° E, 01 Jul 1932, Zakhvatkin, 2♀ (AMNH_PBI 00261497, AMNH_PBI 00261498) (ZISP). Padun' on Verkhnyaya Tunguska River, 56.28333 ° N 101.71666 ° E, 1867, A. Czekanowski, 1♂ (AMNH_PBI 00261297) (ZISP). Kabardino-Balkaria Rep.: Goluboe Lake, Nalchik, 43.48333 ° N 43.6 ° E, 13 Aug 1917, Bartenev, ( Chloanthaceae ), 2♀ (AMNH_PBI 00261492, AMNH_PBI 00261493) (ZISP). Valley of Khuda-Khurt River nr Nalchik, Terskiy Dist., 43.48333 ° N 43.6 ° E, 14 Aug 1917, Bartenev, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261330) (ZISP). Kalmykia Rep.: Yashaltinskoe Lake, Manych Dist., Stavropol Government, 46.33 ° N 42.26 ° E, Aug 1914, B.P. Uvarov, 1♀ (AMNH_PBI 00261504) (ZISP). Karachay-Cherkessia Rep.: Azgek River, Tributary of Teberda River, Kuban' Prov., 43.46666 ° N 41.68333 ° E, 01 Jul 1915 - 02 Jul 1915, L.I. Bogdanova-Katkova, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261172) (ZISP). Karachaevsk [Mikoyan-Shakhar], confluence of Kuban' and Teberda Rivers, 43.76666 ° N 41.9 ° E, 23 Jun 1935, Dyakonov, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261173) (ZISP). Sourse of Daut River, Daut, 43.63 ° N 42.1 ° E, 02 Aug 1935, D'yakonov, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261181) (ZISP). Teberda, 43.45 ° N 41.73333 ° E, 09 Aug 1953, E. Arens, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261306) (ZISP). Karelia Rep.: Petrozavodsk, 61.793 ° N 34.338 ° E, 24 Jul 1905, Jakobson, 1♂ (AMNH_PBI 00261299) (ZISP). Kemerovo Prov.: Berchikul', 55.638 ° N 83.301 ° E, 05 Jul 1898, Plotnikov, 1♂ (AMNH_PBI 00261358), 1♀ (AMNH_PBI 00261496) (ZISP). Fusion of Balyksa River and Magazy River, E of Tom' River, 53.66666 ° N 88.08333 ° E, 1908, Khvorov, 1♀ (AMNH_PBI 00235277) (ZISP). Krasnodar Terr.: Adagum, 6.4 km [6 verst] from Krymskaya, 45.08333 ° N 37.71666 ° E, 06 Jul 1906 - 08 Jul 1906, Vorobyev, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261179) (ZISP). Gelendzhik, 44.5731 ° N 38.0747 ° E, 13 Jul 1956, K. Arnoldi, ( Chloanthaceae ), 2♀ (AMNH_PBI 00261178, AMNH_PBI 00261304) (ZISP). Georgiyevskoye, nr Tuapse, 44.1 ° N 39.08333 ° E, 03 Jun 1910, Smirnov, 2♀ (AMNH_PBI 00261313, AMNH_PBI 00261314) (ZISP). Krasnaya Polyana, Western Caucasus, 43.68333 ° N 40.2 ° E, 1907, A. N. Kiritshenko, ( Chloanthaceae ), 2♂ (AMNH_PBI 00261182, AMNH_PBI 00261183) (ZISP); 04 Jun 1907, A. N. Kiritshenko, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261184) (ZISP). Krasnoaleksandrovskiy Vtoroy [Krasnoaleksandrovskaya], 44.01666 ° N 39.33333 ° E, 11 Jun 1910, Smirnov, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261174) (ZISP). Kuban' Prov., 44.9 ° N 40.23333 ° E, 06 Jul 1990, Vorobyev, 1♀ (AMNH_PBI 00261146) (ZISP). Maly Bombak nr Psebay, Southern Kuban', 44.1 ° N 40.76666 ° E, 15 Jul 1909, Filatov, 3♂ (AMNH_PBI 00159643, AMNH_PBI 00225934, AMNH_PBI 00261144), 1♀ (AMNH_PBI 00261145) (ZISP). Markotkh Range nr Gelendzhik, 44.5731 ° N 38.0747 ° E, 15 Jul 1956, Kurcheva, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261303) (ZISP). Slavyansk-na-Kubani [Slavyanskaya], lower course of Kuban river, 45.23333 ° N 38.11666 ° E, 20 Jun 1936, Rysakov, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261175) (ZISP). Leningrad Prov.: Chernaya Lakhta, 59.967 ° N 29.248 ° E, 02 Jun 1897, Bianchi, 1♂ (AMNH_PBI 00261475) (ZISP); 23 Jun 1899, Bianchi, 1♀ (AMNH_PBI 00260914) (ZISP); 08 Aug 1899, Bianchi, 1♀ (AMNH_PBI 00260915) (ZISP). Gorelovo, 59.76666 ° N 30.1 ° E, 21 Jun 1898, Novotortsev, 1♀ (AMNH_PBI 00260917) (ZISP); 13 Jul 1898, Novotortsev, 1♂ (AMNH_PBI 00260998) (ZISP). Islands on Neva River, Shlisselburg distr., 59.946 ° N 31.037 ° E, 28 Jun 1906, Jakobson, 1♀ (AMNH_PBI 00261281) (ZISP). Kaupilovo [Kaypolovo], nr Gorskaya, 60.033 ° N 29.982 ° E, 09 Jul 1897, Birulya, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261346) (ZISP). Kharlamova Gora, 58.97361 ° N 29.32972 ° E, 18 Jul 1896, Bikhner, 1♀ (AMNH_PBI 00261253) (ZISP); 21 Jul 1896, Bikhner, 1♀ (AMNH_PBI 00261254) (ZISP); 14 Jun 1898 - 18 Jun 1898, Bikhner, 1♂ (AMNH_PBI 00261030) (ZISP); 19 Jun 1898 - 22 Jun 1898, Bikhner, 1♂ (AMNH_PBI 00261028), 2♀ (AMNH_PBI 00261249, AMNH_PBI 00261250) (ZISP); 23 Jun 1898, Bikhner, 1♂ (AMNH_PBI 00261031) (ZISP); 23 Jun 1898 - 26 Jun 1898, Bikhner, 1♂ (AMNH_PBI 00261032), 1♀ (AMNH_PBI 00261255) (ZISP); 10 Jul 1898, Bikhner, 2♂ (AMNH_PBI 00261029, AMNH_PBI 00265629) (ZISP); 15 Jul 1898, Bikhner, 1♀ (AMNH_PBI 00261251) (ZISP); 17 Jul 1898, Bikhner, 1♀ (AMNH_PBI 00261252) (ZISP). Krupeli, 58.8 ° N 29.949 ° E, 21 Jun 1898, Mazarakiy, 1♀ (AMNH_PBI 00261205) (ZISP); 03 Aug 1898, Mazarakiy, 1♀ (AMNH_PBI 00261204) (ZISP). Lakhta, Petrograd Prov., 59.93333 ° N 34.98333 ° E, 15 Jul 1935, Sidorskiy, 1♀ (AMNH_PBI 00261308) (ZISP). Lebyazhye, 59.93333 ° N 29.41666 ° E, 05 Jun 1897, Bianchi, 1♂ (AMNH_PBI 00261476), 1♀ (AMNH_PBI 00260864) (ZISP); 09 Jun 1897, Bianchi, 1♀ (AMNH_PBI 00260865) (ZISP); 11 Jun 1897, Bianchi, 1♂ (AMNH_PBI 00261467), 2♀ (AMNH_PBI 00260869, AMNH_PBI 00260870) (ZISP); 12 Jun 1897, Bianchi, 2♂ (AMNH_PBI 00260466, AMNH_PBI 00260468), 2♀ (AMNH_PBI 00260871, AMNH_PBI 00260872) (ZISP); 16 Jun 1897, Bianchi, 4♂ (AMNH_PBI 00260469 through AMNH_PBI 00260471), 4♀ (AMNH_PBI 00260873 through AMNH_PBI 00260876), 1 juvenille (AMNH_PBI 00260987) (ZISP); 18 Jun 1897, Bianchi, 1♂ (AMNH_PBI 00260472) (ZISP); 22 Jun 1897, Bianchi, 2♂ (AMNH_PBI 00260476, AMNH_PBI 00260477), 2♀ (AMNH_PBI 00260882, AMNH_PBI 00260883) (ZISP); 25 Jun 1897, Bianchi, 1♀ (AMNH_PBI 00260886) (ZISP); 28 Jun 1897, Bianchi, 1♂ (AMNH_PBI 00260986), 1♀ (AMNH_PBI 00260889) (ZISP); 29 Jun 1897, Bianchi, 1♂ (AMNH_PBI 00268033) (ZISP); 26 Jul 1897, Bianchi, 1♀ (AMNH_PBI 00260888) (ZISP); 09 Aug 1897, Bianchi, 1♀ (AMNH_PBI 00260866) (ZISP); 20 Jun 1898, Bianchi, 3♀ (AMNH_PBI 00252582, AMNH_PBI 00260880 through AMNH_PBI 00260881) (ZISP); 25 Jun 1898, Novotortsev, 1♀ (AMNH_PBI 00260891) (ZISP); 27 Jun 1898, Bianchi, 3♂ (AMNH_PBI 00260473 through AMNH_PBI 00260475), 3♀ (AMNH_PBI 00260877 through AMNH_PBI 00260879) (ZISP); 28 Jun 1898, Novotortsev, 1♀ (AMNH_PBI 00260890) (ZISP); 02 Jul 1899, Bianchi, 4♂ (AMNH_PBI 00260461 through AMNH_PBI 00260464), 3♀ (AMNH_PBI 00260861 through AMNH_PBI 00260863) (ZISP); 26 Jul 1900, Bianchi, 1♂ (AMNH_PBI 00268032), 1♀ (AMNH_PBI 00260887) (ZISP); 23 Jun 1907, Bianchi, 1♂ (AMNH_PBI 00260985), 2♀ (AMNH_PBI 00260884, AMNH_PBI 00260885) (ZISP); 10 Jul 1909, Bianchi, 1♂ (AMNH_PBI 00260465), 2♀ (AMNH_PBI 00260867, AMNH_PBI 00260868) (ZISP). Lesnoy: St.-Petersburg, park nr Academy of Forestry Engineering, 59.982 ° N 30.36 ° E, 30 Jun 1898, Bianchi, 2♂ (AMNH_PBI 00261288, AMNH_PBI 00261289) (ZISP). Ligovo, Baltiyskaya Railway, 60.2167 ° N 31.8 ° E, 07 Jul 1898, Novotortsev, 2♀ (AMNH_PBI 00260916, AMNH_PBI 00260919) (ZISP). Lobanovo, 59.817 ° N 30.932 ° E, 20 Jun 1905, A. Vlasov, 1♂ (AMNH_PBI 00261294) (ZISP); 06 Jul 1905, A. Vlasov, 1♀ (AMNH_PBI 00261292) (ZISP); 26 Jul 1908, A. Vlasov, 1♀ (AMNH_PBI 00261293) (ZISP). Log, 58.85 ° N 30.36666 ° E, 18 Jun 1918 - 01 Jul 1918, Jakobson, 1♂ (AMNH_PBI 00260673), 4♀ (AMNH_PBI 00261277 through AMNH_PBI 00261280) (ZISP). Lopukhinka, 59.733 ° N 29.401 ° E, 04 Jul 1894, Bianchi, 2♀ (AMNH_PBI 00260906, AMNH_PBI 00260907) (ZISP); 07 Jul 1894, Bianchi, 2♀ (AMNH_PBI 00260904, AMNH_PBI 00260905) (ZISP); 12 Jul 1894, Bianchi, 1♂ (AMNH_PBI 00260997), 1♀ (AMNH_PBI 00260913) (ZISP); 16 Jul 1894, Bianchi, 1♀ (AMNH_PBI 00260912) (ZISP); 18 Jul 1894, Bianchi, 1♂ (AMNH_PBI 00260996) (ZISP); 31 Jul 1894, Bianchi, 1♂ (AMNH_PBI 00260995), 1♀ (AMNH_PBI 00260910) (ZISP); 01 Aug 1894, Bianchi, 1♀ (AMNH_PBI 00260911) (ZISP); 02 Aug 1894, Bianchi, 2♀ (AMNH_PBI 00260908, AMNH_PBI 00260909) (ZISP). Luga, 58.73333 ° N 29.85 ° E, Jun 1915 - Jul 1915, Kudryavtsev, 1♀ (AMNH_PBI 00261290) (ZISP). Lyazevo nr Siverskaya Station, Varshavskaya Railroad, 59.33333 ° N 30.05 ° E, 27 Jun 1898, Markov, 1♀ (AMNH_PBI 00261203) (ZISP); 09 Jul 1898, N. Kuznetsov, 2♂ (AMNH_PBI 00261041, AMNH_PBI 00261042), 2♀ (AMNH_PBI 00261269, AMNH_PBI 00261270) (ZISP); 24 Jul 1898, Markov, 1♂ (AMNH_PBI 00260980) (ZISP); 31 Jul 1898, N. Kuznetsov, 2♀ (AMNH_PBI 00261271, AMNH_PBI 00261272) (ZISP). Novyy Petergof [Petrodvorets], 59.86666 ° N 29.91666 ° E, 02 Aug 1895, Chekini, 1♀ (AMNH_PBI 00260920) (ZISP). Preobrazhenka railway station, direction of Varshavskaya railway, 59.649 ° N 28.201 ° E, 07 Jul 1898, Pleske, 1♀ (AMNH_PBI 00260976) (ZISP). Romanovka nr Kingisepp [Yamburg], 59.36666 ° N 28.6 ° E, 19 Jun 1905, Barovsky, 1♀ (AMNH_PBI 00261287) (ZISP); 23 Jun 1905, Barovsky, 1♀ (AMNH_PBI 00260979) (ZISP); 02 Jul 1905, Barovsky, 1♀ (AMNH_PBI 00260978) (ZISP); 13 Jul 1905, Barovsky, 1♀ (AMNH_PBI 00260977) (ZISP). Roshchino [Rayvola], Finland Railway, 60.25 ° N 29.61666 ° E, 04 Jul 1896, Prikhodko, 1♂ (AMNH_PBI 00261284) (ZISP); 07 Jul 1896, Prikhodko, 1♀ (AMNH_PBI 00261285) (ZISP). Sablino, 59.61667 ° N 30.73333 ° E, 41 m, 07 Jul 1921, Bianchi, 1♀ (AMNH_PBI 00260918) (ZISP). Serezhino, 59.43 ° N 28.32 ° E, 17 Jun 1895, Bianchi, 3♀ (AMNH_PBI 00260847 through AMNH_PBI 00260849) (ZISP); 20 Jun 1895, Bianchi, 1♀ (AMNH_PBI 00260852) (ZISP); 01 Jul 1895, Bianchi, 1♂ (AMNH_PBI 00260438) (ZISP); 03 Jul 1895, Bianchi, 1♂ (AMNH_PBI 00260440), 1♀ (AMNH_PBI 00260441) (ZISP); 04 Jul 1895, Bianchi, 2♂ (AMNH_PBI 00260442, AMNH_PBI 00260443), 1♀ (AMNH_PBI 00260836) (ZISP); 09 Jul 1895, Bianchi, 3♀ (AMNH_PBI 00260842 through AMNH_PBI 00260844) (ZISP); 23 Jul 1895, Bianchi, 1♀ (AMNH_PBI 00260856) (ZISP); 28 Jul 1895, Bianchi, 1♀ (AMNH_PBI 00260858) (ZISP); 10 Aug 1895, Bianchi, 2♂ (AMNH_PBI 00260459, AMNH_PBI 00260460) (ZISP); 15 Jun 1896, Bianchi, 3♂ (AMNH_PBI 00260445, AMNH_PBI 00260446), 1♀ (AMNH_PBI 00260846) (ZISP); 16 Jun 1896, Bianchi, 2♂ (AMNH_PBI 00260447) (ZISP); 17 Jun 1896, Bianchi, 5♂ (AMNH_PBI 00260448 through AMNH_PBI 00260451) (ZISP); 19 Jun 1896, Bianchi, 1♀ (AMNH_PBI 00260851) (ZISP); 22 Jun 1896, Bianchi, 2♂ (AMNH_PBI 00260452, AMNH_PBI 00260453), 3♀ (AMNH_PBI 00260853 through AMNH_PBI 00260855) (ZISP); 25 Jun 1896, Bianchi, 1♂ (AMNH_PBI 00260456) (ZISP); 26 Jun 1896, Bianchi, 2♂ (AMNH_PBI 00260454, AMNH_PBI 00260455), 1♀ (AMNH_PBI 00260857) (ZISP); 05 Jul 1896, Bianchi, 3♀ (AMNH_PBI 00260837 through AMNH_PBI 00260839) (ZISP); 06 Jul 1896, Bianchi, 1♂ (AMNH_PBI 00260444), 2♀ (AMNH_PBI 00260840, AMNH_PBI 00260841) (ZISP); 09 Jul 1896, Bianchi, 2♂ (AMNH_PBI 00260457, AMNH_PBI 00260458), 2♀ (AMNH_PBI 00260859, AMNH_PBI 00260860) (ZISP); 02 Aug 1896, Bianchi, 1♂ (AMNH_PBI 00260439) (ZISP). Serezhino, 59.43333 ° N 28.31667 ° E, 14 m, 14 Jun 1896, Bianchi, 1♀ (AMNH_PBI 00260845) (ZISP). Shapki [Khimozi] nr Gatchina, 59.6 ° N 31.15 ° E, 10 Aug 1940, A.K. Lukyanovich, 1♀ (AMNH_PBI 00261286) (ZISP). St.-Petersburg, 59.935 ° N 30.31 ° E, V. Jakovlev coll., 1♂ (AMNH_PBI 00261350), 1♀ (AMNH_PBI 00261471) (ZISP). Ushkovo [Tyurisevya], 60.18333 ° N 29.61666 ° E, 20 Jul 1915, Kizeritskiy, 1♀ (AMNH_PBI 00261309) (ZISP). Zelenogorsk [Terioki], 60.201 ° N 29.702 ° E, 13 Jul 1899, Novotortsev, 1♂ (AMNH_PBI 00258643), 1♀ (AMNH_PBI 00260982) (ZISP). Moscow Prov.: Belye Kolodezi, Kolomna distr., 54.55 ° N 38.43 ° E, 08 Jun 1903, G.A. Kozhevnikov, 1♀ (AMNH_PBI 00261510) (ZISP). Porech'ye, 55.7 ° N 35.55 ° E, 09 Jun 1903, Bianchi, 1♂ (AMNH_PBI 00261298) (ZISP). Nizhegorod Prov.: Zhukovka vill., Bor distr., 56.052 ° N 44.673 ° E, 11 Jul 1979, Khrynova, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261168) (ZISP); 31 Jul 1979, Khrynova, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261169) (ZISP). North Osetia Rep.: Dargavs nr Vladikavkaz, 42.83333 ° N 44.41666 ° E, 24 Jul 1925, A. N. Kiritshenko, 4♂ (AMNH_PBI 00260999 through AMNH_PBI 00261002), 6♀ (AMNH_PBI 00260921 through AMNH_PBI 00260926) (ZISP). Dzhimara, 42.819 ° N 44.367 ° E, 26 Jul 1925, A. N. Kiritshenko, 1♂ (AMNH_PBI 00261004) (ZISP). Egochkal nr Dzheyrakh, 42.78333 ° N 44.68333 ° E, 24 Jul 1927, A. N. Kiritshenko, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261329) (ZISP). Koban nr Vladikavkaz, 42.9 ° N 44.46666 ° E, 20 Jul 1925, A. N. Kiritshenko, 1♀ (AMNH_PBI 00260930) (ZISP); 21 Jul 1925, A. N. Kiritshenko, 1♂ (AMNH_PBI 00261003), 3♀ (AMNH_PBI 00260927 through AMNH_PBI 00260929) (ZISP). Pass between Verkhnyaya Kora and Ardon Rivers, 42.82306 ° N 43.91583 ° E, 01 Aug 1925, A. N. Kiritshenko, 1♂ (AMNH_PBI 00261355), 1♀ (AMNH_PBI 00261482) (ZISP). Verkhniy Kakadur [Kakadur] nr Vladikavkaz, 42.83333 ° N 44.38333 ° E, 27 Jul 1925, A. N. Kiritshenko, 2♀ (AMNH_PBI 00260931, AMNH_PBI 00260932) (ZISP). Vladikavkaz, 43.01666 ° N 44.66666 ° E, 11 Jul 1925, A. N. Kiritshenko, 1♂ (AMNH_PBI 00261005) (ZISP). Novgorod Prov.: Tigoda River, 59.286 ° N 31.571 ° E, 14 Jul 1903, Semenov, 1♀ (AMNH_PBI 00261193) (ZISP); 23 Jul 1909, Semenov, 1♀ (AMNH_PBI 00261194) (ZISP). Voron'ya Gora, 58.5 ° N 32.465 ° E, 05 Jun 1898, Shmidt, 1♂ (AMNH_PBI 00261295) (ZISP). Penza Prov.: E Gorodishche, Penza Gov., 53.26666 ° N 45.68333 ° E, 04 Jul 1864, Chekanovskiy, 1♀ (AMNH_PBI 00261199) (ZISP). unknown, 26 Jun 1864, A. Czekanowski, 1♂ (AMNH_PBI 00261200) (ZISP). Perm Terr.: Perm, 58.01666 ° N 56.3 ° E, 1925, Lubischev, 1♀ (AMNH_PBI 00260981) (ZISP). Pskov Prov.: Glebova Gorka, 58.61666 ° N 29.01666 ° E, 08 Jul 1861, Moravits, 1♀ (AMNH_PBI 00250722) ( Chloanthaceae ), 1♂ (AMNH_PBI 00261348) (ZISP); 16 Jul 1961, Moravits, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261344) (ZISP). Oparino, Novorzhevsk, distr. Pskov, 57.0333 ° N 29.3333 ° E, 11 Jul 1915, N. Kuznetsov, 1♀ (AMNH_PBI 00261317) (ZISP). Shchepets [former Yaz'vo Volost', Gdov Government], 58.75 ° N 28.2 ° E, 29 Jun 1901, Ivanov, 1♂ (AMNH_PBI 00261198) (ZISP); 12 Jul 1901, Ivanov, 1♂ (AMNH_PBI 00250452) (ZISP); 16 Jul 1901, Ivanov, 2♀ (AMNH_PBI 00261195, AMNH_PBI 00261196) (ZISP); 20 Jul 1901, Ivanov, 1♀ (AMNH_PBI 00261197) (ZISP). Ryazan Prov.: Gremyachka, 53.91666 ° N 39.5 ° E, 21 Jul 1908, Semenov, 1♀ (AMNH_PBI 00261154) (ZISP); 03 Jul 1909, Semenov, 1♂ (AMNH_PBI 00261160), 2♀ (AMNH_PBI 00261152, AMNH_PBI 00261153) (ZISP); 22 Jun 1911, Semenov, 1♂ (AMNH_PBI 00261162), 1♀ (AMNH_PBI 00261156) (ZISP); 04 Jul 1911, Semenov, 1♂ (AMNH_PBI 00261161) (ZISP); 02 Jul 1912, Semenov, 1♀ (AMNH_PBI 00261157) (ZISP); 04 Jul 1912, Semenov, 1♂ (AMNH_PBI 00261163), 1♀ (AMNH_PBI 00261158) (ZISP); 14 Jun 1913, Semenov, 1♀ (AMNH_PBI 00261159) (ZISP). Sten'kino, 54.5 ° N 39.5 ° E, 14 Jul 1913, Bianchi, ( Chloanthaceae ), 2♀ (AMNH_PBI 00261170, AMNH_PBI 00261171) (ZISP). Samara Prov.: Alekseevka, Buzuluk Distr., 52.88333 ° N 51.65 ° E, 22 Jun 1909 - 23 Jun 1909, Bakhtin, ( Chloanthaceae ), 1♂ (AMNH_PBI 00264806) (ZISP). Saratov Prov.: Aleksandrov-Gay, Samara Prov., 50.13333 ° N 48.55 ° E, Jun 1912, Neustroev, ( Chloanthaceae ), 1♀ (AMNH_PBI 00227522) (ZISP). Pugachevsk [Nikolaevsk], 52.01666 ° N 48.8 ° E, 25 Jun 1925, Dyakonov, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261301) (ZISP); 01 Jul 1925, Dyakonov, 1♀ (AMNH_PBI 00260983) ( Chloanthaceae ), 1♀ (AMNH_PBI 00261302) (ZISP); 08 Jul 1925, Dyakonov, 1♀ (AMNH_PBI 00260984) (ZISP). Stavropol Terr.: Bryk-Gora, 44.61666 ° N 42.63333 ° E, 07 Jul 1914, B.P. Uvarov, 1♂ (AMNH_PBI 00265156), 1♀ (AMNH_PBI 00253500) ( Chloanthaceae ), 5♂ (AMNH_PBI 00261015 through AMNH_PBI 00261019), 11♀ (AMNH_PBI 00261235 through AMNH_PBI 00261245) (ZISP). Kislovodsk, North Caucasia, 43.88333 ° N 42.71666 ° E, 30 Jul 1908, Lubischev, ( Chloanthaceae ), 1♀ (AMNH_PBI 00250424) (ZISP); 15 Jul 1914, Bartenev, ( Chloanthaceae ), 1♂ (AMNH_PBI 00261307), 1♀ (AMNH_PBI 00261305) (ZISP). Makhach-Aul [Dzelal-Ogly], Caucasus, 44.45 ° N 45.01666 ° E, 16 Aug 1920, Unknown, ( Chloanthaceae ), 2♀ (AMNH_PBI 00261494) (ZISP). Sengileyevskaya [Sengileyevskoye], 45.03333 ° N 41.65 ° E, 26 May 1914, B.P. Uvarov, ( Chloanthaceae ), 2♀ (AMNH_PBI 00261177, AMNH_PBI 00261300) (ZISP). Stavropol, 45.05 ° N 41.95 ° E, 20 Jun 1914, B.P. Uvarov, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261176) (ZISP). Yessentuki, Terskaya Prov., 44.03333 ° N 42.83333 ° E, 14 Jul 1914, Bartenev, ( Chloanthaceae ), 2♀ (AMNH_PBI 00261185, AMNH_PBI 00261186) (ZISP); 29 Jul 1920, Ryabov, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261187) (ZISP). Sverdlovsk Prov.: Ekaterinburg, 56.84638 ° N 60.6075 ° E, 25 Jun 1909, Dyakonov, 1♀ (AMNH_PBI 00261513) (ZISP); 09 Aug 1909, Dyakonov, 1♀ (AMNH_PBI 00261468) (ZISP). Tatarstan Rep.: Kazan', 55.78333 ° N 49.16666 ° E, Eversmann coll, 2♂ (AMNH_PBI 00261351, AMNH_PBI 00261353), 1♀ (AMNH_PBI 00261472) (ZISP). Tuva Rep.: Berezovka, 51.249 ° N 94.333 ° E, 29 Jun 1949, Dyatlova, 1♀ (AMNH_PBI 00261500) (ZISP). Low Course of Sistiq-Khem River, 52.66666 ° N 95.5 ° E, 21 Jul 1897 - 22 Jul 1897, Ostrovskiy, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261347) (ZISP). Vologda Prov.: Nr Vologda, 59.21666 ° N 39.88333 ° E, 07 Jul 1895, Birulya, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261164) (ZISP). Spasskoye-Kurkino vill., 59.36 ° N 39.63 ° E, 19 Jul 1921, L.I. Bogdanova-Katkova, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261166) (ZISP); 29 Jul 1921, L.I. Bogdanova-Katkova, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261165) (ZISP). Tot'ma, 59.96666 ° N 42.75 ° E, 12 Jul 1933, Barovsky, 1♂ (AMNH_PBI 00261147) (ZISP); 15 Jul 1934, Barovsky, 2♀ (AMNH_PBI 00261148, AMNH_PBI 00261149) (ZISP); 26 Jul 1935, Barovsky, 1♀ (AMNH_PBI 00261150) (ZISP); 31 Jul 1935, Barovsky, 1♂ (AMNH_PBI 00225684), 1 juvenille (AMNH_PBI 00261151) (ZISP). Voronezh Prov.: Kamennaya Steppe 11 km S Talovaya, 51.21666 ° N 40.71666 ° E, 24 Jun 1928, Unknown, 1♂ (AMNH_PBI 00261363), 1♀ (AMNH_PBI 00261502) (ZISP); 15 Jun 1935, Stark, 1♀ (AMNH_PBI 00261077) (ZISP); 23 Jun 1935, Stark, 8♂ (AMNH_PBI 00149992, AMNH_PBI 00261081 through AMNH_PBI 00261087), 4♀ (AMNH_PBI 00238060, AMNH_PBI 00261078 through AMNH_PBI 00261080) (ZISP); 08 Jun 1949, Stark, 1♀ (AMNH_PBI 00261076) (ZISP). Ternovka [Ternovskiy sovkhoz], Elan'-Kolenovskiy distr., 49.6 ° N 39.4667 ° E, 19 May 1937, Frolova, 5 juvenille (AMNH_PBI 00261043 through AMNH_PBI 00261047) (ZISP); 07 Jul 1937, Frolova, 1♀ (AMNH_PBI 00150044) (ZISP); 11 Jul 1951, Stark, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261169) (ZISP). Voronezh, 51.6664 ° N 39.17 ° E, 23 Jun 1947, Bodrenkov, 1♂ (AMNH_PBI 00258644) (ZISP). Yaroslavl Prov.: Berditsino, 57.482 ° N 40.1 ° E, 29 Apr 1893, Unknown, 1♀ (AMNH_PBI 00261514) (ZISP); 18 Jun 1896, Unknown, 1♂ (AMNH_PBI 00260674) (ZISP); 24 Jun 1896, Unknown, Salix sp. (Salicaceae) , 1♀ (AMNH_PBI 00261054) (ZISP); 26 Jun 1896, Unknown, 2♂ (AMNH_PBI 00260675) (ZISP); 27 Jun 1896, Unknown, 2♀ (AMNH_PBI 00261050, AMNH_PBI 00261051) (ZISP); 13 Jul 1896, Unknown, 1♀ (AMNH_PBI 00261055) (ZISP). Bykovo, 58.13333 ° N 38.6 ° E, 11 Jul 1895, Unknown, 3♀ (AMNH_PBI 00261052, AMNH_PBI 00261053) (ZISP). Pavlovskoe, 57.9411 ° N 39.6097 ° E, 23 Jun 1891, Unknown, 1♀ (AMNH_PBI 00261515) (ZISP). SPAIN: Caceres: Guadalupe, Caceres, 39.45 ° N 5.32 ° W, 12 May 1952, Gonzalez, 1♂ (AMNH_PBI 00268025) (JR). Catalonia: Alcover, 41.25 ° N 1.16666 ° E, 07 Jun 1981, J. Ribes, 1♂ (AMNH_PBI 00268026), 1♀ (AMNH_PBI 00268026) (JR). Caldas de Montbuy [Caldes de Montbui], 41.64 ° N 2.17 ° E, 12 Jun 1977, J. Ribes, 1♂ (AMNH_PBI 00268027), 2♀ (AMNH_PBI 00268027) (JR). Galicia: Between Granja and Segovia (De la Granja a Segovia), 42.9 ° N 7.46 ° W, 08 Jul 1957, J. Ribes, 2♀ (AMNH_PBI 00231113) (JR). La Rioja: Cabezon de Cameros, Logrono, 42.22 ° N 2.52 ° W, 13 Aug 1978, J. Ribes, 1♀ (AMNH_PBI 00234534) (JR). Madrid: Casa de Campo, 40.42 ° N 3.75 ° W, May 1957, Unknown, 1♂ (AMNH_PBI 00268024) (JR). Valldoreix (Valles Ocid.), 41.45 ° N 2.03 ° W, 24 Jun 1972, J. Ribes, 1♂ (AMNH_PBI 00228062) (JR). TURKEY: Antalya: Altinyaka, 36.55 ° N 30.33333 ° E, 28 Apr 2002, Doguet, 1♀ (AMNH_PBI 00264064) (MATOCQ). Ancient City of Termessos (24 mi N of Antalya), 37.348 ° N 31 ° E, 12 May 1998, Doguet, 2♂ (AMNH_PBI 00263374, AMNH_PBI 00263375) (MATOCQ). Kale, 36.25 ° N 29.98333 ° E, 27 Apr 2002, Doguet, 1♂ (AMNH_PBI 00264059) (MATOCQ). Icel Ili: Camliyayla [Namrun], 37.16666 ° N 34.6 ° E, 26 Jun 1962 - 30 Jun 1962, Seidenstücker, 1♀ (AMNH_PBI 00268054) (ZSM). Isparta: Atabey, 37.95 ° N 30.63333 ° E, 14 May 1998, Doguet, 1♂ (AMNH_PBI 00263371), 2♀ (AMNH_PBI 00264065) (MATOCQ). Mersin: Tarsus, 36.9178 ° N 34.8917 ° E, 17 m, 23 Apr 1955, Seidenstücker, 1♂ (AMNH_PBI 00157874), 2♀ (AMNH_PBI 00157874) (JR) 1♀ (AMNH_PBI 00268052) (ZSM); 04 May 1958, Seidenstücker, 1♀ (AMNH_PBI 00268053) (ZSM). Adana, 16 May 1960, Eckerlein, 1♂ (AMNH_PBI 00150876), 4♀ (AMNH_PBI 00237612, AMNH_PBI 00268071 through AMNH_PBI 00268073) (MHNG). Ankara, 39.91666 ° N 32.85 ° E, 02 Jun 1963 - 03 Jun 1963, Seidenstücker, 10♂ (AMNH_PBI 00227520, AMNH_PBI 00235129, AMNH_PBI 00268035 through AMNH_PBI 00268038, AMNH_PBI 00268040 through AMNH_PBI 00268043) (ZSM). Ankara (Gýlbaschi), 12 Jun 1962, Seidenstücker, 1♀ (AMNH_PBI 00268044) (ZSM); 18 Jun 1965, Seidenstücker, 2♀ (AMNH_PBI 00268045, AMNH_PBI 00268046) (ZSM). Ankara - Cubukbaraji, 13 Jul 1962, Seidenstücker, 1♀ (AMNH_PBI 00225859) (ZSM). Ankara Baraj, 40.23333 ° N 38.91667 ° E, 25 May 1965 - 27 May 1965, Seidenstücker, 2♀ (AMNH_PBI 00268047, AMNH_PBI 00268048) (ZSM); 02 May 1968 - 03 May 1968, Seidenstücker, 2♂ (AMNH_PBI 00268050, AMNH_PBI 00268051) (ZSM); 06 Jun 1968, Seidenstücker, 1♀ (AMNH_PBI 00268049) (ZSM). Kayseri Yilanil-Dagh, 38.42 ° N 35.28 ° E, 03 Jun 1965 - 07 Jun 1965, Seidenstücker, 1♀ (AMNH_PBI 00159273) (ZSM). Posanti [Pozanti] Taurus, 37.415 ° N 34.871 ° E, 09 May 1955 - 15 May 1955, Seidenstücker, 2♀ (AMNH_PBI 00268055, AMNH_PBI 00268056) (ZSM). UKRAINE: Crimea: Agarmish Mt., 44.48333 ° N 34.16666 ° E, A. N. Kiritshenko, 1♀ (AMNH_PBI 00228087) (ZISP); 25 May 1906, V. Jakovlev coll., 1♀ (AMNH_PBI 00261071) (ZISP); 03 Jun 1906, A. N. Kiritshenko, 1♂ (AMNH_PBI 00261074) (ZISP); 07 Jun 1906, V. Jakovlev coll., 1♀ (AMNH_PBI 00261349) (ZISP); 08 Jun 1906, A. N. Kiritshenko, 1♀ (AMNH_PBI 00261073) (ZISP); 11 Jun 1906, A. N. Kiritshenko, 1♂ (AMNH_PBI 00261075) (ZISP); 22 Jun 1906, V. Jakovlev coll., 1♀ (AMNH_PBI 00261072) (ZISP). Ayudag, southern Coast of Crimea, 44.57 ° N 34.33 ° E, 18 Jun 1938, A. N. Kiritshenko, 4♂ (AMNH_PBI 00237923, AMNH_PBI 00261137 through AMNH_PBI 00261139), 2♀ (AMNH_PBI 00228131, AMNH_PBI 00261136) (ZISP). Kerch, 45.33333 ° N 36.45 ° E, 01 Jun 1917, A. N. Kiritshenko, 3♂ (AMNH_PBI 00259797, AMNH_PBI 00261006 through AMNH_PBI 00261007), 2♀ (AMNH_PBI 00261206, AMNH_PBI 00261207) (ZISP). Simferopol near Salgir river, 44.95 ° N 34.08333 ° E, 14 Jul 1924, A. N. Kiritshenko, ( Chloanthaceae ), 1♀ (AMNH_PBI 00261180) (ZISP). Bagovitsa, Khmelnyts'ka distr., 48.6 ° N 26.71666 ° E, 19 Jun 1896, Saynog, 1♀ (AMNH_PBI 00266004) (ZISP). Gribovitsa [Chernyakovo], Ostrozhskiy Uezd, Volynskaya Gov., 50.68 ° N 24.24 ° E, 01 Jul 1900 - 05 Jul 1900, Neklyudov, 3♀ (AMNH_PBI 00261097 through AMNH_PBI 00261099) (ZISP). Kirivograd [Elisavetograd], 48.5042 ° N 32.2631 ° E, 21 Jun 1902, Yatsentkovskiy, 2♀ (AMNH_PBI 00261470) (ZISP). Krasna Mts. Range [Polonina Krasna], Zakarpats'ka Prov., 48.33 ° N 23.83 ° E, 20 Aug 1953, Roshko, 1♀ (AMNH_PBI 00261096) (ZISP). Mekhedovka, Zolotonoshsk uezd, Poltavskaya prov., 49.85 ° N 32.25 ° E, 12 Jun 1916, Kizeritskiy, 1♀ (AMNH_PBI 00261311) (ZISP). Nr Kaniv, Shevchenkvskiy Nature Reserve, 49.75 ° N 31.46666 ° E, 21 Jul 1932, S. I. Medvedev, 1♂ (AMNH_PBI 00261365) (ZISP); 29 Jul 1932, S. I. Medvedev, 1♀ (AMNH_PBI 00261506) (ZISP). Nr Zhitomir, 50.25 ° N 28.65 ° E, 09 Jun 1897, Birulya, 1♀ (AMNH_PBI 00261122), 1 juvenille (AMNH_PBI 00261129) (ZISP); 10 Jun 1897, Birulya, 1♀ (AMNH_PBI 00261123) (ZISP); 13 Jun 1897, Birulya, 1♀ (AMNH_PBI 00261124) (ZISP); 17 Jun 1897, Birulya, 1♀ (AMNH_PBI 00261125) (ZISP); 20 Jun 1897, Birulya, 2♀ (AMNH_PBI 00261126, AMNH_PBI 00261128) (ZISP); 21 Jun 1897, Birulya, 1♀ (AMNH_PBI 00261127) (ZISP). Nyrkovo-Vyemka Steppe, Donetsk Terr., 48.7 ° N 38.22 ° E, 20 Jun 1937, L.V. Arnoldi, 1♂ (AMNH_PBI 00261316), 1♀ (AMNH_PBI 00261315) (ZISP). Odessa, Kuyalnitskiy Liman, 46.55 ° N 30.73333 ° E, 06 May 1920, A. N. Kiritshenko, 1♀ (AMNH_PBI 00261101) (ZISP). Pechiski [former Proskurov Uezd of Podolsk Government], 49.53333 ° N 27.1 ° E, 10 Jul 1909, Elachich, 4♂ (AMNH_PBI 00257913, AMNH_PBI 00261140 through AMNH_PBI 00261142) (ZISP); 08 Jul 1910, Elachich, 1♀ (AMNH_PBI 00225969) (ZISP); 11 Jul 1910, Elachich, 1♀ (AMNH_PBI 00261143) (ZISP). Poltava, 49.58333 ° N 34.55 ° E, 15 Jul 1925, A.K. Lukyanovich, 1♂ (AMNH_PBI 00261312), 1♀ (AMNH_PBI 00261310) (ZISP). Provalye, 48.16666 ° N 39.83333 ° E, 16 Jun 1908, Troitskiy, 1♂ (AMNH_PBI 00261320) (ZISP); 21 Jun 1947, S. I. Medvedev, 2♂ (AMNH_PBI 00261322, AMNH_PBI 00261323), 1♀ (AMNH_PBI 00261321) (ZISP). Shutova River nr Yuzhny Bug River, Voznesensk Distr.,, 47.577 ° N 31.331 ° E, 15 Jul 1922, A. N. Kiritshenko, 2♂ (AMNH_PBI 00261133, AMNH_PBI 00261134), 3♀ (AMNH_PBI 00261130 through AMNH_PBI 00261132) (ZISP). Slavuta, 50.28333 ° N 26.86666 ° E, 01 Jul 1911, Zaytsev, 1♂ (AMNH_PBI 00259800) (ZISP); 08 Jul 1911, F. A. Zaytsev, 5♀ (AMNH_PBI 00224843, AMNH_PBI 00261102 through AMNH_PBI 00261105) (ZISP); 12 Jul 1911, F. A. Zaytsev, 2♂ (AMNH_PBI 00261108, AMNH_PBI 00261109), 2♀ (AMNH_PBI 00261106, AMNH_PBI 00261107) (ZISP). Velikie Korovintsy [Korovitsy], nr Mikhalenki st., 49.983 ° N 28.283 ° E, Jun 1948 - Jul 1948, Gulinov, 1♀ (AMNH_PBI 00261100) (ZISP). Verkhovka [former Mahilyow uezd], 48.9 ° N 27.65 ° E, 24 May 1901, Chekini, 1♂ (AMNH_PBI 00261135) (ZISP); 02 Jun 1901, Chekini, 1♀ (AMNH_PBI 00261192) (ZISP). Yelanets, NE of Voznesensk, 47.68333 ° N 31.85 ° E, 10 Jul 1923, A. N. Kiritshenko, 4♀ (AMNH_PBI 00261067 through AMNH_PBI 00261070) (ZISP). YUGOSLAVIA: Serbia: Topcidersko Brdo [Topchider] nr Belgrad, 44.76666 ° N 20.45 ° E, 07 Jun 1927, Martino, 2♂ (AMNH_PBI 00261035, AMNH_PBI 00261036) (ZISP); 08 Jun 1927, Martino, 1♂ (AMNH_PBI 00261037) (ZISP); 10 Jun 1927 - 15 Jun 1927, Martino, 1♂ (AMNH_PBI 00261040) (ZISP); 15 Jun 1927, Martino, 1♂ (AMNH_PBI 00261038) (ZISP); 09 Jun 1928 - 10 Jun 1928, Martino, 1♀ (AMNH_PBI 00261039) (ZISP).