Mesabolivar González-Sponga, 1998

Huber, Bernhard A., 2018, The South American spider genera Mesabolivar and Carapoia (Araneae, Pholcidae): new species and a framework for redrawing generic limits, Zootaxa 4395 (1), pp. 1-178 : 6-7

Mesabolivar González-Sponga, 1998

Mesabolivari [sic] González-Sponga, 1998: 27. Type species by original designation: M. pseudoblechroscelis González- Sponga, 1998.

Mesabolivar: Huber 2000: 189 . Huber 2015: 5. Machado 2007: 50.

Autana González-Sponga, 2011b: 40 . Type species by original designation: A. autanensis González-Sponga, 2011 . Synonymized in Huber et al. 2014a: 419.

Caruaya González-Sponga, 2011b: 40 . Type species by original designation: C. anseriformis González-Sponga, 2011 . Synonymized in Huber et al. 2014a: 418.

Kaliana Huber, 2000: 271 . Type species by original designation: K. yuruani Huber, 2000 . Synonymized in Astrin et al. 2007: 23.

Rioparaguanus González-Sponga, 2005: 104. Type species by original designation: R. spinosus González-Sponga, 2005 . Synonymized in Huber et al. 2014a: 418.

Teuia Huber, 2000: 313 . Type species by original designation: Teuia beckeri Huber, 2000 . New synonymy.

Justification of synonymy. The genus Teuia was originally proposed for a single species ( T. beckeri ) that has a highly unusual procursus (partly wrapped around the bulbal process; Huber 2000: figs 1259–1260) but lacks an epigynal pocket (Huber 2000: fig. 1263) and contiguous male cheliceral apophyses (Huber 2000: fig. 1262). The new species M. sepitus described below shares the distinctive morphology of the procursus and was collected only 130 km from the type locality of Teuia beckeri . However, M. sepitus has an epigynal pocket and contiguous male cheliceral apophyses. Molecular data (Eberle et al., unpublished data; see Appendices 1–2) also support a position of this new species within the Southern clade of Mesabolivar . Teuia is thus synonymized with Mesabolivar , and T. beckeri is transferred accordingly: Mesabolivar beckeri (Huber, 2000) , comb. n.

Note, however, that the name Teuia is an available name in case Mesabolivar is split (e.g. for the “Southern clade” in Appendices 1-2). Another genus that might be revalidated in such a context is Kaliana Huber, 2000 . As indicated above (Introduction), I do not implement such drastic formal changes here for two reasons: (1) current evidence is rather limited as far as taxon sampling and support values are concerned; (2) the gain of such formal changes is dubious if the diagnosability of the resulting genera is not improved.

Notes. For recent updates on generic diagnosis and description, see Huber (2015). Most of the new species described below fit this generic concept that dates back to Huber (2000) and was further developed in Machado (2007). The two key features are (1) an epigynum with median groove or pocket (e.g., Figs 20 , 112 , 223 ), and (2) male cheliceral apophyses with a pair of contiguous frontal apophyses (e.g., Figs 18 , 118, 215). Most species that do not fit this concept are part of what is below informally described as the ‘ difficilis group’. Several species in this group of litter-dwelling species do not share any of the diagnostic characters. However, molecular data consistently place them among Mesabolivar . The same is true for individual representatives of other species groups proposed below, for example in the togatus group. In addition, it is now fairly evident that the two key features are shared by certain representatives of the closely related genus Carapoia (e.g., C. lutea , C. tapajos , C. maculata ). In its current scope, a morphological diagnosis that covers all species of Mesabolivar is probably impossible.

Figures

FIGURES18–25. Mesabolivar acrensissp.n. (18–22; ZFMK Ar 18942–43) andM.maraba sp. n.(23–25, ZFMK Ar 18948). 18–19. Male chelicerae, frontal and lateral views. 20–21, 23–24. Epigyna, ventral and lateral views. 22, 25. Cleared female genitalia, dorsal views.Abbreviations:ep, epigynal pocket; pp, pore plate; v, ‘valve’. Scale lines:0.5.

FIGURES 104–114. Left bulbal processes, dorsal and prolatero-ventral views (104–111), epigynum, ventral and lateral views (112–113), andcleared female genitalia,dorsal view (114). 104–105.Mesabolivar macushi sp. n. (ZFMK Ar 18962).106–107. M. aurantiacus (Mello-Leitão, 1930) (ZFMK Ar 18963). 108–109. M.cyaneus (Taczanowski, 1874) (ZFMK Ar 18989). 110– 114.M.spinosus (González-Sponga, 2005) (MIZA MAGS 1176). Scale lines: 0.5 (all bubal processes at same scale).

FIGURES 223–226. Mesabolivar madalena sp. n. (223–224, ZFMK Ar 19071) and M. inmanis sp. n. (225–226, ZFMK Ar 19072). 223, 225. Epigyna, ventral views. 224, 226. Cleared female genitalia, dorsal views (arrow points at V-shaped sclerite). Scale lines:0.5.