Canthon (Glaphyrocanthon) quadriguttatus (Olivier, 1789)

Chamorro, William, Marin-Armijos, Diego, senjo, Angelico & Vaz-De-Mello, Fernando Z., 2019, Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records, ZooKeys 826, pp. 1-343 : 54

Canthon (Glaphyrocanthon) quadriguttatus (Olivier, 1789) Plate 13D

Scarabeus quadriguttatus Olivier, 1789: 173 (original description. Type locality: Cayenne, Surinam).

Copris quadriguttatus : Olivier 1790: 178 (new combination, redescription).

Ateuchus bidens Fabricius, 1801: 62 (transferred to the genus Ateuchus Fabricius, 1801); Harold 1868d: 123 (synonym of C. quadriguttatus Oliv.).

Choeridium elegans Castelnau, 1840: 83 (transferred to the genus Choeridium Serville, 1825; redescription); Harold 1868d: 123 (synonym of C. quadriguttatus Oliv.).

Canthon quadriguttatus : Harold 1868d: 123 (transferred to the genus Canthon Hoffmannsegg 1817, redescription); Gemminger and Harold 1869: 993 (list, distribution); Gillet 1911a: 32 (complete list of species); Schmidt 1920: 125 (comment, list of species); Medina et al. 2001: 136 (cited for Colombia); Ratcliffe et al. 2015: 195 (cited for Peru).

Glaphyrocanthon quadriguttatus : Pereira and Martínez 1956a: 132 (new combination, catalog of species); Martínez et al. 1964: 12 (characters in key); Vulcano and Pereira 1964: 664 (catalog of species); Vulcano and Pereira 1967: 560 (characters in key).

Canthon (Glaphyrocanthon) quadriguttatus : Halffter and Martínez 1977: 80 (list of species); Rivera-Cervantes and Halffter 1999: 44 (characters in key); Vaz-de-Mello 2000: 191 (cited for Brazil, cited as C. quadriguttatum ); Medina et al. 2003: 64 (distribution); Krajcik 2012: 64 (complete list of species); Boilly and Vaz-de-Mello 2013: 112 (figure 36); Chamorro et al. 2018: 93 (cited for Ecuador).

Copris obliquatus Voet, 1806: 47 (original description).

Canthon obliquatus : Schmidt 1920: 125 (comment, list of species); Schmidt 1922: 78 (distribution); Boucomont 1928c: 2 (distribution [ quadriguttatus Olivier], cited for Guyana); Balthasar 1939d: 205 (characters in key); Balthasar 1941: 342 (cited for Peru); 1951: 327 (cited for Peru); Halffter and Martínez 1977: 80 (synonym of Canthon quadriguttatus , (Olivier, 1789)); Vaz-de-Mello and Cupello 2018b: 49 (cited as junior objective synonym of Canthon (Glaphyrocanthon) quadriguttatus (Olivier, 1789), comment).

Type specimens.

Scarabeus quadriguttatus Olivier, 1789. Not found.

Ateuchus bidens Fabricius, 1801. One syntype examined deposited at the ZMUC (ex coll. E Harold and R Oberthur). Lectotype to be designated in a future work on this species group.

Choeridium elegans Castelnau, 1840. Type material not examined.

Canthon obliquatus Schmidt, 1920. Two syntypes examined deposited at the NHRS. Lectotype to be designated in a future work on this species group.

Distribution.

Brazil, Colombia, Ecuador, Peru, Guyana, and Surinam.

Records examined.

ORELLANA: El Dorado plataforma Guarango, 300 m (1 specimen MUTPL); El Dorado, plataforma Pitalala 1, 325 m (1 specimen MUTPL); road Auca Tiguino-Ñemenguno, Parque Nacional Yasuní, 350 m (1 specimen MUTPL). PASTAZA: Bosque Protector Oglán Alto, 545 m (2 specimens MUTPL); Pandanuque, 420 m (1 specimen CEMT). SUCUMBÍOS: Aucayacu Río El Eno, 16 km de Lago Agrio, 295 m (1 specimen MUTPL); El Dorado de Cascales, Pozo Cristal 1, 425 m (1 specimen MUTPL); Tarapoa Campo Marian, plataforma Fanny 5 (1 specimen MUTPL); Tarapoa Campo Marian, plataforma Fanny 18B60, 245 m (1 specimen MUTPL). TUNGURAHUA: Baños (1 specimen MUTPL). ZAMORA CHINCHIPE: Tundayme campamento Mirador, Las Maravillas, 1060 m (3 specimens MUTPL); Tundayme campamento Mirador, San Marcos, 900 m (1 specimen MUTPL).

Temporal data.

Collected in March, May, July, June, August, October, November, and December.

Remarks.

Inhabits the lowland evergreen forests and the foothill evergreen forests of the Amazon region from 245-1060 m a.s.l. Collected in aerial fruit traps, with canopy fogging methods and pitfall traps baited with carrion and human feces.