Phanerotoma vanharteni, Achterberg, 2021

Achterberg, Cornelis van, 2021, The tribe Phanerotomini (Hymenoptera, Braconidae, Cheloninae) of the Arabian Peninsula, with special reference to the United Arab Emirates and Yemen, ZooKeys 1014, pp. 1-118 : 1

Phanerotoma vanharteni sp. nov. Figs 374-376 , 377-386

Type material.

Holotype, ♀ (RMNH), ♀, " Yemen: Al Kowd (8136), ix.2003, light trap, A. v. Harten & S. Al Haruri, RMNH’03”. Paratypes: 2♀: Idem, v.-vi.2000; 1♀: Idem, x.2000; 1♀: Idem, 27-31.vii.2001; 1 ♂: "Yemen (6628), Ta’izz, light trap, vi.2002, A. van Harten & A.R. Al Yarimi, RMNH"; 2♀: " United Arab Emirates, NARC near Sweihan (1473), light trap, 28.iii.-2.iv.2005, 24°24'N, 55°26'E, A. v. Harten, RMNH’05”.

Diagnosis.

Apical antennal segments of ♀ short and distinctly moniliform (Figs 374 , 382 , 386 ), sixth segment from apex narrowed basally and subapically widened (Fig. 382 ); upper condyle of mandible near lower level of eyes (Fig. 383 ); temple narrow in lateral view (median width of temple 0.4-0.5 × width of eye; Fig. 385 ); width of clypeus 0.9 × minimum width of face (Fig. 383 ); height of eye in anterior view 1.3 × minimum width of face; face densely sculptured, rather matt and dorsally often with fine median carina (Fig. 383 ), but sometimes superficially sculptured, without median carina and shiny; hind tibia subbasally usually partly dark brown; inner tooth of mandible medium-sized (Fig. 376 ); vein r of fore wing 0.8-1.1 × vein 3-SR; hypopygium of ♀ straight apically in lateral view (Figs 375 , 380 ), without up curved apical triangle or spine-like protuberance.

Description.

Female, holotype, length of body (excluding ovipositor) 3.5 mm; antenna 2.5 mm; fore wing 2.5 mm; visible part of ovipositor sheath 0.5 mm, only apically with erect setae.

Head. Width 1.6 × median length in anterior view and part of head above eye in lateral view 0.25 × height of eye (Fig. 385 ); antenna with 23 segments and as long as fore wing, segments near apical quarter of antenna longer than wide, somewhat serrate and widened subapically because of small round protuberances and nine apical segments moniliform (Figs 382 , 386 ) and apical segment with spine, third, fourth and penultimate segments 3.4, 2.8 and 1.1 × longer than wide in lateral view, respectively; area of stemmaticum coriaceous; OOL: diameter of posterior ocellus: POL = 7: 3: 5; length of eye 5.4 × temple in dorsal view (Fig. 384 ); frons rugose laterally, rugulose medially and with satin sheen, without median carina posteriorly; vertex rugose but posteriorly striate and with satin sheen; temple densely striate and rather matt, its median width 0.4 × width of eye in lateral view; face densely sculptured, rather matt and dorsally with fine median carina; clypeus largely very finely coriaceous, with satin sheen and its width 0.9 × minimum width of face, intertentorial distance 3.0 × minimum distance between clypeus and eye and medio-ventrally with three minute lobes (Fig. 383 ); eye large, strongly convex, in anterior view its height 1.3 × minimum width of face; upper condyle of mandible near lower level of eyes (Fig. 383 ); malar space rugose, with satin sheen and its length 0.5 × basal width of mandible; lower tooth of mandible 0.5 × as long as apical tooth (Fig. 376 ).

Mesosoma (Figs 374 , 378 ). Length 1.5 × its width in lateral view; side of pronotum granulate dorsally, rugose medially and remainder mostly smooth; propleuron posteriorly evenly convex and protruding towards medial sulcus; mesosternum densely granulate and matt; mesoscutum densely rugulose; scutellum flat, densely granulate but smooth posteriorly and with satin sheen; notauli not indicated; scutellar sulcus wide and with six carinae (Fig. 378 ); metanotum with median carina anteriorly, flat posteriorly and narrowly serrate posteriorly; propodeum granulate-rugulose anteriorly and remainder coarsely rugose-reticulate, transverse carina distinct, without median carina, antero-dorsal face long and latero-posteriorly slightly tuberculate. Wings. Fore wing 2.6 × longer than its maximum width; length of 1-R1 1.4 × as long as pterostigma; distance between wing apex and vein 1-R1 0.2 × length of vein 1-R1; r issued distinctly beyond middle of pterostigma, nearly linearly connected to 3-SR and 0.9 × 3-SR; 2-SR straight and distally slightly converging to posterior margin of pterostigma (Fig. 377 ); SR1 straight; 2-SR+M present, m-cu antefurcal; parastigma large; 1-CU1 0.5 × as long as vein 2-CU1, cu-a slightly inclivous and 0.8 × 1-CU1; r:3-SR:SR1 = 10:11:47; 2-SR:3-SR:r-m = 20:11:7; r-m reclivous; 2-M slightly curved (Fig. 377 ). Hind wing: M+CU:1-M:1r-m = 23:19:10. Legs. Hind femur rather dull, 3.5 × as long as wide and widened submedially; hind tibia wide medially (Fig. 381 ); middle tibia with small ivory blister; inner spur of middle tibia 0.55 × its basitarsus; hind coxa largely superficially granulated and rather shiny.

Metasoma (Figs 379 , 380 ). Elliptical in dorsal view, 1.9 × as long as wide and 1.3 × as long as mesosoma; first and second tergites coarsely longitudinally rugose; metasomal sutures medium-sized and deep; third tergite distinctly convex medially, 1.7 × longer than second tergite and laterally curved, in lateral view rather convex (Fig. 380 ), largely densely reticulate-rugulose and with satin sheen, lateral lamella narrow laterally and posteriorly, not protruding latero-apically and medio-apically truncate; ovipositor sheath parallel-sided and rather narrow apically, its visible part 0.15 × as long as fore wing and 0.3 × metasomal carapace and only apically with medium-sized setae; hypopygium setose and acute apically, without up curved triangle or spine apically (Fig. 380 ).

Colour. Yellowish brown; apical antennal segments apically and apex of ovipositor sheath brown; stemmaticum dark brown; parastigma largely, veins 1-CU1, base of 2-CU1, cu-a, r, 2-SR, 3-SR, SR1 and 2-M of fore wing dark brown, other veins (including vein 1-M), and pterostigma basally and apically, pale yellowish; apical half of pterostigma partly dark brown; clypeus, mandible (except dark brown teeth), palpi, pronotal side, tegulum, mesoscutum medio-posteriorly, legs (but hind femur rather brownish except basally and hind tibia subbasally and apically dark brown), first and second tergites and metasoma baso-ventrally pale yellowish; humeral plate brownish; wing membrane below veins 1-&2-CU1 and below pterostigma brownish and remainder largely subhyaline.

Male.

Similar to female (including shape of hind femur and tibia); antennal segments slenderer and weakly serrate and with minor subapical protuberances; median width of temple 0.55 × width of eye in lateral view.

Variations.

Length of fore wing of ♀ 3.0-3.8 mm, of ♂ 2.8 mm; median width of temple 0.4-0.5 × width of eye in lateral view; dark part of hind tibia dark brown or brown; vein r of fore wing linear with vein 3-SR or rather angled, 0.8-1.1 × vein 3-SR; face often densely sculptured, rather matt or with satin sheen and dorsally often with fine median carina (Fig. 383 ), but female from UAE has face superficially sculptured, without median carina and shiny.

Biology.

Unknown.

Distribution.

United Arab Emirates, Yemen.

Etymology.

Named in honour of Tony van Harten (editor of the series 'Arthropod Fauna of the UAE’) for his extraordinary efforts to make extensive collections of insects and species in hardly investigated habitats.

Figures

Figures 374 - 376. Phanerotoma vanharteni van Achterberg, sp. nov., ♀, holotype 374 habitus lateral 375 apical half of metasoma lateral 376 mandible ventral.

Figures 377 - 386. Phanerotoma vanharteni van Achterberg, sp. nov., ♀, holotype 377 fore wing 378 mesosoma dorsal 379 first-third metasomal tergites dorsal 380 metasoma lateral 381 hind leg lateral 382 apical third of antenna lateral 383 head anterior 384 head dorsal 385 head lateral 386 antenna lateral.