species, Morley, 1913

Rousse, Pascal & Noort, Simon Van, 2014, A review of the Afrotropical Rhyssinae (Hymenoptera: Ichneumonidae) with the descriptions of five new species, European Journal of Taxonomy 91, pp. 1-42 : 4-11

Key to species

1. Fore wing with areolet closed, triangular and strongly petiolate (A); body orange and dark brown, wings dark brown (B); tropical Africa ………………………… Megarhyssa babaulti Seyrig, 1937

- Fore wing with areolet open (a); body and wings colorations variable but usually wings hyaline and body yellow interspersed with black markings (b) …………………………………2 ( Epirhyssa ) 2. Face smooth to sparsely punctate (A-B), without transverse rugosities …………………………… 3 - Face transversely rugulose punctate or transversely striate (a-b) ……………………………………5

3. Wings hyaline, fore wing slightly infuscate apically (A); Madagascar …… E. migratoria Seyrig, 1932

- Wings with dark patterns, yellowish to black (a, b); tropical Africa ………………………………4

4. Wings blackish with subapical lighter patch (A); epicnemial carina reaching level of mesopleural pit (B) ………………………………………………………………………… E. villemantae sp. nov.

- Wings yellowish with apex blackish and an additional subapical transverse blackish band on fore wing (a);epicnemial carina very short,hardly extending onto mesopleuron(b)……… E.maynei Benoit, 1952

5. Metasoma with large apico-lateral yellow spots on tergites 1–5 (A); clypeus without median subapical tubercle (B); body and ovipositor large to very large (B> 25mm; OT> 6) …… E. overlaeti Seyrig, 1937

- Metasoma usually uniformly colored (a); clypeus with a distinct subapical tubercle (b); body and ovipositor smaller (B <25mm; OT <6) ……………………………………………………………6

6. Anterior sclerotized part of sternite 1 with a pair of submedian tubercles, tubercles also present on sternites 2–4 (A); general coloration dark testaceous to black interspersed with pale yellow (B) …… ………………………………………………………………………………… E. brianfisheri sp. nov.

- Sternite 1 without a pair of submedian tubercles, tubercles present on sternites 2–4 only (a); general coloration pale to striking yellow with isolated black markings (b) …………………………………7 7. Apex of subtegular ridge concave, laterally flanged (A) ……………………………………………8 - Apex of subtegular ridge convex, without lateral flange (a) ……………………………………9

8. Clypeus punctate (A); tergite 1 of female slender, more than 1.4x longer than apically wide (B) ……… ………………………………………………………………………………… E. gavinbroadi sp. nov.

- Clypeus almost smooth, hardly sculptured (a); tergite 1 of female stout, less than 1.4x longer than wide (b) …………………………………………………………………… E. uelensis Benoit, 1951

9. Tergite 2 densely punctate over its entire surface and tergite 1 more than 1.5x longer than apically wide (A); general coloration pale yellow (B); Southern Africa ……………………… E. shaka sp. nov.

- Tergite 2 smooth to superficially punctate laterally (a), if ambiguous (some E. tombeaodiba ) then tergite 1 stouter, less than 1.5x longer than apically wide; general coloration bright yellow (b); tropical Africa ………………………………………………………………………………………10

10. Fore wing with 2m–cu basal to rs–m (A); mesosternum black (B) ……… E. leroyi Benoit, 1951

- Fore wing with 2m–cu apical to rs–m (a); mesosternum yellow (b) ………………………11

11. Clypeus shallowly longitudinally strigose (A); frons with two medial carinae converging in front of median ocellus (B); tergite 1 of female stout, less than 1.4x longer than apically wide (C) ………… …………………………………………………………………………… E. tombeaodiba sp. nov.

- Clypeus smooth (a); frons with two submedian carinae, diverging on upper frons toward lateral ocelli (b); tergite 1 of female slender, more than 1.4x longer than apically wide (c) ………………………………………………………………………… E. ghesquierei Benoit, 1951

Figures

1. Fore wing with areolet closed, triangular and strongly petiolate (A); body orange and dark brown, wings dark brown (B); tropical Africa ………………………… Megarhyssa babaulti Seyrig, 1937

- Fore wing with areolet open (a); body and wings colorations variable but usually wings hyaline and body yellow interspersed with black markings (b) …………………………………2 ( Epirhyssa ) 2. Face smooth to sparsely punctate (A-B), without transverse rugosities …………………………… 3 - Face transversely rugulose punctate or transversely striate (a-b) ……………………………………5

3. Wings hyaline, fore wing slightly infuscate apically (A); Madagascar …… E. migratoria Seyrig, 1932

- Wings with dark patterns, yellowish to black (a, b); tropical Africa ………………………………4

4. Wings blackish with subapical lighter patch (A); epicnemial carina reaching level of mesopleural pit (B) ………………………………………………………………………… E. villemantae sp. nov.

- Wings yellowish with apex blackish and an additional subapical transverse blackish band on fore wing (a);epicnemial carina very short,hardly extending onto mesopleuron(b)……… E.maynei Benoit, 1952

5. Metasoma with large apico-lateral yellow spots on tergites 1–5 (A); clypeus without median subapical tubercle (B); body and ovipositor large to very large (B> 25mm; OT> 6) …… E. overlaeti Seyrig, 1937

- Metasoma usually uniformly colored (a); clypeus with a distinct subapical tubercle (b); body and ovipositor smaller (B <25mm; OT <6) ……………………………………………………………6

6. Anterior sclerotized part of sternite 1 with a pair of submedian tubercles, tubercles also present on sternites 2–4 (A); general coloration dark testaceous to black interspersed with pale yellow (B) …… ………………………………………………………………………………… E. brianfisheri sp. nov.

- Sternite 1 without a pair of submedian tubercles, tubercles present on sternites 2–4 only (a); general coloration pale to striking yellow with isolated black markings (b) …………………………………7 7. Apex of subtegular ridge concave, laterally flanged (A) ……………………………………………8 - Apex of subtegular ridge convex, without lateral flange (a) ……………………………………9

8. Clypeus punctate (A); tergite 1 of female slender, more than 1.4x longer than apically wide (B) ……… ………………………………………………………………………………… E. gavinbroadi sp. nov.

- Clypeus almost smooth, hardly sculptured (a); tergite 1 of female stout, less than 1.4x longer than wide (b) …………………………………………………………………… E. uelensis Benoit, 1951

9. Tergite 2 densely punctate over its entire surface and tergite 1 more than 1.5x longer than apically wide (A); general coloration pale yellow (B); Southern Africa ……………………… E. shaka sp. nov.

- Tergite 2 smooth to superficially punctate laterally (a), if ambiguous (some E. tombeaodiba ) then tergite 1 stouter, less than 1.5x longer than apically wide; general coloration bright yellow (b); tropical Africa ………………………………………………………………………………………10

10. Fore wing with 2m–cu basal to rs–m (A); mesosternum black (B) ……… E. leroyi Benoit, 1951

- Fore wing with 2m–cu apical to rs–m (a); mesosternum yellow (b) ………………………11

11. Clypeus shallowly longitudinally strigose (A); frons with two medial carinae converging in front of median ocellus (B); tergite 1 of female stout, less than 1.4x longer than apically wide (C) ………… …………………………………………………………………………… E. tombeaodiba sp. nov.

- Clypeus smooth (a); frons with two submedian carinae, diverging on upper frons toward lateral ocelli (b); tergite 1 of female slender, more than 1.4x longer than apically wide (c) ………………………………………………………………………… E. ghesquierei Benoit, 1951