Thymbra Linnaeus (1753: 569)

Bräuchler, Christian, 2018, Delimitation and revision of the genus Thymbra (Lamiaceae), Phytotaxa 369 (1), pp. 15-27 : 16-17

Thymbra Linnaeus (1753: 569)

Type (designated by Green 1929): Thymbra spicata Linnaeus (1753: 569) .

= Coridothymus Reichenbach (1857: 160) ≡ Thymus subgen. Coridothymus (Rchb.f.) Borbás (1890: 48) ≡ Thymus sect. Coridothymus (Rchb.f.) Boissier (1879: 560) .

Type: Coridothymus capitatus ( Linnaeus 1753: 568) Reichenbach (1857: 161) ≡ Satureja capitata Linnaeus (1753: 568)

= Euhesperida Brullo & Furnari (1979: 434) .

Type: Euhesperida thymbrifolia Brullo & Furnari (1979: 434) .

= Satureja Linnaeus (1753: 568) et sensu Harley et al. (2004: 239).

Description

Subshrubs, 10–60 cm, aromatic; branches ascending, erect or intricate; young leaves in axillary fascicles, leaves entire, sometimes with few cilia at margin, obovate, elliptic, lanceolate or linear, conduplicate to caniculate or flat, sessile, sometimes slightly succulent, punctate by sessile to shortly stalked globular glands, secondary veins tiny and not visible, glabrous to densely villose; inflorescence a terminal spiciform or capituliform thyrse, condensed to lax, composed of congested verticillasters; bracts leaf like or not, oblong-ovate to ovate-lanceolate or lanceolate, sometimes coloured; bracteoles not leaf like, elliptic to lanceolate, acute or mucronate; calyx 3–6.5 mm long, 2-lipped (sometimes appearing almost actinomorphic due small dimensions and dense indumentum), 5-lobed (3/2), cylindrical to curved, with two +/-conspicuous lateral keels (produced by the veins ending in the lateral lobes of the upper calyx lip), more or less dorsiventrally compressed, scarcely accrescent, 12–23-nerved, lobes never spreading or following the tube outline, either subequal, triangular to acicular, or unequal with posterior lobes short, triangular, anterior lobes lanceolate, upcurved, throat sometimes constricted, never wide, bearded or not; corolla 2-lipped, 4-lobed (1/3), 4–12 mm long, cream or almost white to purple, posterior lobe +/-rounded, straight, emarginate, anterior lip with lobes suborbicular, median lobe downcurved; stames 4, inserted in middle of tube, filaments curved, exserted, thecae divaricate, confluent; stigma-lobes equal to subequal; disc entire; mericarps ovoid, +/-smooth. 2n = 30; seven species, Mediterranean, and in adjacent areas (W: Portugal; E: Anatolia, Iraq and Iran to N Arabian Peninsula).

Thymbra calostachya GREECE. E Crete, 1 km W of Monikapsa , rocks close to the sea, 10 m, 15 June 1990, Ulrich s.n. (M) ; Crete, Sitia, W Kapsa Monastery , 31 March 2007, Tillich 5231 (M, TUM) ; ibid., 01 April 2007, Tillich 5265 (M, TUM)

T. capitata CULTIVATED. [origin: GREECE. Crete, Bräuchler 2526] Botanical Garden Munich , 03 June 2005, Bräuchler 3403 (TUM); [origin SPAIN] Palmengarten Frankfurt , 25 September 2003, Bräuchler 2519 ( TUM) ; GREECE. Crete, Prov. Rethymnon 35°11‘44.0“N 24°24‘21.1“E, 116 m, 26 March 2004, Bräuchler 2526 ( TUM) ; Peloponnes, Taygetos , 15 July 2007, Kattari SK 07/ Pel 046 ( TUM) ; ITALY. Sardinia, 39°03‘00.0“N 8°21‘30.0“E, 81 m, 11 June 1999, Bräuchler 97 ( TUM) ; MALTA. Meimberg 05 ( TUM) ,

T. linearifolia LIBYA. Wadi Belbarabides , 09 May 1974, Brullo & Furnari s.n. ( TUM) ; Wadi el-Chresci (Tocra), 21 May 1981, Brullo & Furnari s.n. ( TUM)

T. nabateorum SAUDI ARABIA. Off the Dula-Tabuk road, 10 km NE of Ruafah, 4.500 ft., 16 May 1994, Collenette 9059 (E00102242) ; 72 km S of Tayma , Madinah road, 17 April 1991, Collenette 7814 (E) ; JORDAN.

12 km SSW of Petra (type locality), 11 May 1995, Danin s.n. ( TUM; HUJ980701 ) ; Edom , Ras Raibid, 640 m, 03 October 1988, Künne 3368 ( TUM) ; 1.5 km SW of Taiyiba , 1000 m, 28 September 1993, Künne 93-123-02 ( TUM) ; Siq Qseir al Anajil ( NW Taiyiba), 1100 m, 30 September 1995, Künne 95- 122-01 ( TUM) ; Edom , El-Hesma , 22 km N. of Queira, Nubian sandstone, about 800 m, 30 March 1936, Eig, Zohary & Feinbrun s.n. (K000193589, paratype of Satureja thymbrifolia )

T. sintenisii subsp. isaurica TURKEY. Antalya, Alanya, 36°45´N, 32°28´E, 1290 m, 18 July 1998, Güner et al. 12628 (E00075637)

T. spicata subsp. spicata CULTIVATED. [origin TURKEY. Taurus, Erben s.n.] Botanical Garden Munich, 23 June 2008, Bräuchler 4548 (TUM); ibid., 26 July 2007, Bräuchler 4197 ; LEBANON. Above Matara on the road to reserve, 33°39‘49.3“N 35°38‘06.4“E, 1025 m, 24 July 2010, Bräuchler 5207 ( TUM) ; TURKEY. Antalya, Alanya , 36°41‘25.6“N 31°53‘26.0“E, 522 m, 12 August 2007, Bräuchler 4420 ( TUM) ; Antalya, Kemer , 36°40‘1.16“N, 30°32‘25.62“E, 40 m, 11 August 2007, Bräuchler 4161 ( TUM) ; Karabük, Safranbolu , 41°13‘53.3“N 32°40‘21.5“E, 363 m, 04 August 2007, Bräuchler, Dirmenci &

Yildiz 4344 (TUM)

T. thymbrifolia ISRAEL: Judean Desert , 11 km S of Jericho, 29 April 1994, Danin s.n. ( TUM) ; Judean Desert , Near Khirbet el Mird Hill, 02 April 1932, Eig & Feinbrun s.n. ( HUJ 315080 )