Stenus

Puthz, Volker, 2015, Neotropische Stenus-Arten mit seitlich gerandetem Abdomen und gelappten Tarsen (Coleoptera, Staphylinidae) 344. Beitrag zur Kenntnis der Steninen, Linzer biologische Beiträge 47 (2), pp. 1601-1727 : 1691-1699

Preliminary key to the neotropic Stenus species with margined abdomen

and bilobed tarsi (exclusively S. holmbergii LYNCH ARRIBALZÁGA )

E = edeagus; S = sternite, T= tergite

1 Body with long, erect, conspicuous pubescence (figs 1-6).................................................2

- Body with recumbent pubescence.......................................................................................3

2 Head broader than elytra, elytra as long as broad with a very small, indistinct reddish spot.: Unknown.: S8 (fig. 5). 4.5-5.3mm (FB2.4-2.5mm). Ecuador............................ ............................................................................................................. longepilosus nov.sp.

- Head narrower than elytra, elytra broader than long with a distinct reddish spot in posterior half (about as in fig. 137).: Unknown.: S8 (about as in fig. 298). 4.0- 4.7mm (FB 2.3mm). Venezuela........................................................... setosiventris nov.sp.

3 Brachypterous.....................................................................................................................4

- Macropterous....................................................................................................................12

4 Elytra with reddish spots.....................................................................................................5

- Elytra black.........................................................................................................................6

5 Median portion of frons densely punctate, elytral spot indistinct, abdominal tergites reticulate.: E (fig. 204).: S8 (fig. 216). 4.0-5.0mm (FB 2.0-2.3mm). Mexico.............. ........................................................................................................... chimalpopoca nov.sp.

- Median portion of frons broadly impunctate, elytral spot distinct, very large (fig. 130), abdominal tergites not reticulate.: E (fig. 203).: S8 (fig. 214). 4.2- 5.0mm (FB 2.3-2.4mm). Bolivia..................................................................................... chasqui nov.sp.

6 Interstices of tergite 7 reticulate ..........................................................................................7

- Interstices of tergite 7 smooth...........................................................................................11

7 Punctuation of abdomen coarse and dense, punctures of tergite 3 much larger than cross section of antennal segment 3....................................................................................8

- Punctuation of tergite 3 less coarse and less dense, punctures of tergite 3 about as large as cross section of antennal segment 3..............................................................................10

8 Larger species, FB>2.3mm................................................................................................9

- Smaller species, FB 2.0-2.1mm.: E (figs 210, 232),: S8 (fig. 215). 3.7-4.4mm. Mexico......................................................................................................... centeotl nov.sp.

9 Punctuation of tergite 7 sparser (and finer), interstices much larger than diameter of punctures.: E (fig. 314). 4.2 mm (FB 2,3mm). Mexico....................... ometochtli nov.sp.

- Punctuation of tergite 7 denser (and coarser), interstices about as large as punctures.: E (figs 366, 377).: S8 (fig. 357). 4.7-5.5mm (FB 2.3-2.4mm). Panama............................ ............................................................................................................. trapezivestis nov.sp.

10 Larger, FB  2,2mm.: E (fig. 218).: T8 (fig. 35), S8 (fig. 283), Valvifer (fig. 114). 3.8-4.7mm (FB 2.2mm). Brazil.................................................................. mrazi L. BENICK

- Smaller, FB  2,1mm,: E (figs 267, 301, 302),: T8 (fig. 34), S8 (fig. 33). 3.7- 4.5mm (FB 2.0-2.1mm). Brazil................................................................ juliae HROMÁDKA

11 Larger, FB 2.4mm. S3 (fig. 9).: E (fig. 191, 192),: S8 (fig. 199). 4.6-5.1mm. Brazil....................................................................................................... blandus L. BENICK

- Smaller, FB 1,9-2,1mm.: E (fig. 266, 299).: S8 (fig. 274). 4.0-4.5mm(FB 1.9- 2.1mm). Brazil............................................................................................ irritator nov.sp.

12 Frons very broad, HW: DE <1.40 (e. g. fig. 47)................................................................13

- Frons less broad, HW: DE>1.50 (e. g. figs 48-52)...........................................................18

13 Elytra black.......................................................................................................................14

- Elytra with reddish spots...................................................................................................16

14 Head much narrower than elytra (HW: EW 0.86) (fig. 47). Antennae sexual dimorph, 4 th tarsal segment very narrow (fig. 23)............................................................................15

- Head about as broad as elytra (HW: EW 1,00).: E (fig. 303).: S8 (fig. 275). 3.5- 4.1mm (FB 2.0-2.1mm). Suriname, French Guiana................................. lancifrons nov.sp.

15 Slightly shining, elytral punctuation at places obliquely confluent.: E (figs 174, 179). 3.0-3.6mm (F 1.9mm). Ecuador..................................................... nigrificatus PUTHZ

- Nearly matt, elytral punctuation not confluent. E (fig. 180). 3.2-3.8mm (FB 1.8- 1.9mm). Peru....................................................................................... obscurefactus PUTHZ

16 Each elytron with two reddish sports (fig. 123).: E (fig. 157). 3.8-4.3mm. S8 (as in S. aratorius ) (FB 2.1-2.2mm). Venezuela....................................................... arator PUTHZ

- Each elytron with one reddish spot (figs 124, 125)...........................................................17

17 Elytra (fig. 124).: E (fig. 186, 200).: S8 (fig. 196). 3.2-3.9mm (FB 1.7-1.9mm). Ecuador. .................................................................................................... aratorius nov.sp.

- Elytra (fig. 125).: Unknown.: as in S. aratorius . 3.6-4.0mm (FB 2.0-2.1mm). Costa Rica................................................................................................... aratrifer nov.sp.

18 Elytra black (if there seems to be an indistinct spot follow #67)......................................19

- Elytra with reddish spots (if indistinct, these species are keyed twice).............................68

19 Elytra ± densely punctate..................................................................................................20

- Elytra finely and sparsely punctate, interstices twice and more as large as punctures (fig. 69); head much broader than elytra.: E (fig. 270). 3.0-3.7mm (FB 1.6-1.7mm). West Indies: Grenada................................................................................. lucens CAMERON

20 Whole abdomen reticulate reticulation may be faint or missing on tergite 3 and 4).........21 - Only tergites 7 (and 8) reticulate OR abdomen completely shiny.....................................28

21 Larger, FB> 3,4mm ...........................................................................................................22

- Smaller, FB <3,0mm.........................................................................................................23

22 Sculpture of elytra densely rugose/confluent (fig. 67).: E (fig. 254, 262).: S8 (fig. 244). 6.8-8.0mm (FB 3.6-4.0mm). Costa Rica............................................. hidalgo nov.sp.

- Sculpture of elytra not rugose/confluent (fig. 68).: E (fig. 164).: S8 rounded at posterior margin. 6.5-8.0mm (FB 3.5-3.7mm). Brazil............................... colosseus PUTHZ

23 Median portion of frons feebly elevated, nearly flat, densely punctate. Smaller, FB 2.1mm.: E (fig. 173).: S8 (fig. 326). 4.0-4.8mm (FB 2.0-2.1mm). French Guiana.... ........................................................................................................................ [ rothi PUTHZ ]

- Median portion of frons distinctly elevated, posterior portion or completely smooth. Larger, FB 2.2mm...........................................................................................................24

24 Interstices on tergite 7 larger, twice as large as punctures................................................25

- Interstices on tergite 7 smaller, about as large as punctures or smaller.............................27

25 Larger, FB 2.7-2.8mm. Punctuation of pronotum and elytra rugose/confluent.Head less broad (HW: EW 0.93).....................................................................................................26

- Smaller, FB 2.3-2.6mm. Punctuation of pronotum and elytra dense, the punctures mostly discrete, not confluent. Head broader (HW: EW 0.97) ........................................... ................................................................................................ [ novoteutonicus WENDELER ]

26 Elytral sculpture less rugose (figs 74, 75).: S8 (fig. 30), E (figs 345-349, 351-354).: S8 (fig. 341). 4.9-6.3mm (FB 2.6-2.9mm). Mexico, Honduras, Costa Rica, Panama...... .................................................................................................................. scabrosus SHARP

- Elytral sculpture more distinctly rugose (fig. 76).: S8 (fig. 31), E (figs 365, 378).: S8 (fig.358). 5.2-7.0mm (FB 2.7-2.9mm). Mexico, Guatemala, Honduras........................... ........................................................................................................... trochanterinus SHARP

27 Abdominal punctuation coarse and dense, interstices on tergite 4 as large as or smaller than half diameter of punctures. Paratergites 5 impunctate. Elytral sculpture rugose.: Metatibae without an additional tooth, E (figs 234-236).: S8 (fig. 230). 4.1-4.7mm (FB 2.1-2.4mm). Mexico.......................................................................... emily HROMÁDKA

- Abdominal punctuation less coarse and less dense, interstices on tergite 4 larger than half diameter of punctures. Paratergites 5 punctate. Punctures of elytra distinctly discrete.: Metatibiae with an additional tooth (fig. 10), E (fig. 220).: Unknown. 4.6mm (FB 2.2mm). Brazil..................................................................... condei WENDELER

28 Pronotum broader than long or as broad as long...............................................................29

- Pronotum longer than broad..............................................................................................33

29 Paratergites 5 punctate, tergite 10 concave at posterior margin........................................30

- Paratergites 5 impunctate, tergite 10 simple......................................................................32

30 Punctures of elytra distinctly delimited, paratergites 5 as broad as antennal segment 2.: T10 (fig. 29), E fig. 208),: S8 (fig. 213), Valvifer (fig. 115). 4.0-4.7mm (FB 2.1- 2.3mm). Argentina................................................................................. bruchi BERNHAUER

- Punctures of elytra often confluent, Paratergite 5 narrower than antennal segment 2.: E (fig. 331). 3.8-5.0mm (FB 2.0-2.1mm). Brazil......................................... perssoni PUTHZ

31 Interstices on tergite 7 at most as large as punctures.........................................................32

- Interstices on tergite 7 distinctly larger than punctures.: E (figs 369, 370).: S8 (fig. 356). 4.5-5.8mm (FB 2.2-2.3mm). Brazil..................................... theresae HROMÁDKA

32 Larger, FB  2,4mm.: E (figs 238, 239).: S8 (fig. 26). 4.8-5.3mm. Paraguay............... ................................................................................................................. falinianus nov.sp.

- FB  2,1mm.: E (figs 168, 240).: S8 (fig. 229). 3.9-4.4mm (FB 2.1mm). French Guiana............................................................................................................ grocae PUTHZ

33 Base of abdominal tergites with remarkably dense whitish pubescence (fig. 8).: E (figs 315, 230).: S8 (fig. 295). 3.8-4.5mm (FB 2.0-2.2mm). Panama, Colombia, Brazil, Ecuador, Peru, Bolivia............................................................... ornativentris SHARP

- Base of abdominal tergites moderately densely pubescent...............................................34

34 Antennae very slender, segment 10 2.5x as long as broad (fig. 7).: Unknown.: S8 (fig. 325). 3.0-4.5mm (FB 2.2mm). French Guiana. ........................... procericornis nov.sp.

- Antennae less slender, segment 10 at most twice as long as broad...................................35

35 Elytra broader (EW: EL 1.10).........................................................................................36

- Elytra less broad (EW: EL <1.10).....................................................................................42

36 Sternum 9 apicolaterally acute (fig. 39)............................................................................37

- Sternum 9 apicolaterally ± distinctly serrate (figs 37, 40, 43-46).....................................38

37 Tergite 7 reticulate. Frons deeply concave, median portion nearly flat, punctate.: E (figs 25, 171).: S8 (fig. 323). 4.0-4.7mm (FB 2.1-2.3mm). Honduras, Guatemala, El Salvador, Mexico Costa Rica.................................................................... penicillus nov.sp.

- Tergite 7 not reticulate.: E (figs 371, 379).: S8 (fig. 372). 3.5-4.5mm (FB 2.0- 2.1mm). Ecuador, Peru.............................................................................. yupanqui nov.sp.

38 Larger: FB 2.3mm...........................................................................................................39

- Smaller: FB 2.3mm.........................................................................................................40

39: E (fig. 162).: S8 (fig. 211). 5.0-5.7mm (FB 2.5-2.7mm). Brazil................................. ........................................................................................................ brasilianus BERNHAUER

-: E (figs. 307, 308).: (fig. 284). 4.4-5.5mm (FB 2.3-2.6mm). Brazil............................. ................................................................................................... novoteutonicus WENDELER

40 Punctuation of tergite 7 less dense, interstices predominantly as large as punctures or larger.................................................................................................................................41

- Punctuation of tergite 7 denser, interstices mostly smaller than punctures.: E (fig. 190).: S8 (fig. 189). 3.4-4.5mm (FB 1.8-2.1mm). Brazil, French Guiana, Suriname, Peru.................................................................................................. bechyneae HROMÁDKA

41 Abdomen more cylindrical.: E (fig. 222).: S8 (fig. 246). Brazil, Paraguay, Argentina, Chile................................................................................. endosquameus PUTHZ

- Abdomen more ellipsoid,: E (figs 350, 374).: Unknown. 4.0-4.5mm (FB 2.2mm). Brazil................................................................................................... spathulipenis nov.sp.

42 Tergite 7 reticulate............................................................................................................43

- Tergite 7 not reticulate......................................................................................................48

43 Punctuation of the lateral portions of frons moderately sparse, interstices often as large as punctures.......................................................................................................................44

- Punctuation of lateral portions of frons dense, interstices mostly smaller than half diameter of punctures........................................................................................................45

44 Abdominal punctuation very fine and sparse, punctures on tergite 7 smaller than eye facets near medial eye margins; punctuation of elytra sparser, interstices often as large as punctures or larger.: E (figs 258, 259, 268).: S8 (fig. 280). 4.7-5.5mm (FB 2.4- 2.7mm). Ecuador...................................................................................... janae HROMÁDKA

- punctuation less fine and less sparse, punctures on tergite 7 as large as eye facets near medial eye margins; punctuation of elytra denser, interstices mostly smaller than half diameter of punctures.: E (figs 257, 265).: S8 (fig. 248). 3.9-5.0mm (FB 2.4- 2.5mm). Panama.................................................................................... intolerabilis nov.sp.

45 Larger, FB  2,4mm ...........................................................................................................47

- Smaller, FB 2.3mm.........................................................................................................46

46 FB 2.1-2.2mm, abdominal punctuation denser, interstices of tergite 7 at most as large as punctures.: S8 (fig. 28), E (fig. 173).: S8 (fig. 326). 4.0-4.8mm (FB 2.0- 2.1mm). French Guiana...................................................................................... rothi PUTHZ

- Larger, FB  2,3mm, abdominal punctuation less dense, interstices of tergite 7 much larger than punctures.: E (fig. 364). 4.5-5.0mm (FB 2.3mm). Costa Rica........................ ............................................................................................................ toripennis L. BENICK

47 Larger (FB 2.5-2.7mm), head less broad (HW: EW 0.92).: Distance of preapical tooth of metatibae about as large as size of tooth. E (fig. 162).: S8 (fig. 211). 5.0- 5.7mm (FB 2.5-2.7mm). Brazil....................................................... brasilianus BERNHAUER

- Smaller (FB 2.0-2.5mm).: Distance of preapical tooth of metatibae at least twice as large as size of tooth.: E (figs 165, 221).: S8 (fig. 228). 4.0- 5.0mm (FB 2.0- 2.5mm). Costa Rica, Nicaragua, Honduras, Belize, Guatemala, Mexico.............................. ........................................................................................................... costipennis L. BENICK

48 Lateral portions of frons densely punctate, interstices mostly smaller than half diameter of punctures.......................................................................................................................49

- Lateral portions of frons less densely punctate, interstices often larger than half diameter of punctures.: E (fig. 272).: S8 as in fig. 249). 4.2-5.0mm (FB 2.4- 2.5mm). Ecuador....................................................................................... ignoratus nov.sp.

49 Punctuation of tergite 7 sparser, interstices mostly as large or distinctly larger than punctures...........................................................................................................................50

- Punctuation of tergite 7 denser, interstices distinctly smaller than punctures...................63

50 Head less broad (HW: EW 0.97).....................................................................................51

- Head broader (HW: EW 1,00)........................................................................................53

51 Elytral sculpture coarser, punctures about as large as medial cross section of antennal segment 2..........................................................................................................................52

- sculpture less coarse, punctures about as large as apical cross section of antennal segment 2..........................................................................................................................54

52 Sculpture of pronotum in posterior half transversely confluent.: E (fig 223, 224).: S8 as in fig. 228. 4.3-5.0mm (FB 2.1-2.3mm). Costa Rica.................... denticulifer nov.sp.

- Sculpture of pronotum in posterior half rugose but not transversely confluent.................53

53: E (figs 165, 201).: S8 (fig. 228). 4.0-5.0mm (FB 2.0-2.5mm). Costa Rica, Nicaragua, Honduras, Belize, Guatemala, Mexico............................ costipennis L. BENICK

-: E (fig. 364).: S8 as in fig. 228. 4.5-5.0mm (FB 2.3mm). CostaRica ........................... ............................................................................................................ toripennis L. BENICK

54 Larger, FB 2.4-2.6mm.......................................................................................................55

- Smaller, FB 2.4mm.........................................................................................................56

55 Punctuation of tergite 7 finer and sparser, interstices twice and more as large as punctures.: E (fig. 162).: S8 (fig. 211). 5.0-5.7mm (FB 2.5-2.7mm). Brazil................ ........................................................................................................ brasilianus BERNHAUER

- Punctuation of tergite 7 coarser and denser, interstices about as large as punctures.: S8 (fig. 27), E (figs 177, 181, 285-292).: S8 (fig. 281). 5.4-6.0mm (FB 2.4- 2.6mm). Panama, French Guiana, Costa Rica, Honduras, Belize, Mexico............................ ............................................................................................................... longiventris SHARP

56 Elytral punctuation very dense, interstices distinctly smaller than half diameter of punctures...........................................................................................................................57

- Elytral punctuation less dense, interstices often as large or larger than diameter of punctures...........................................................................................................................62

57 Sternum 9 apicolaterally serrate.: E (fig. 335).: Unknown. 3.8-4.3mm (FB 2.0- 2.1mm). Brazil.............................................................................................. pedator SHARP

- Sternum 9 apicolaterally acute..........................................................................................58

58 Partergites coarsely and densely punctate.: E (figs 362, 376). 3.7-4.3mm (FB 2.1- 2.2mm). Venezuela................................................................................ subnotatus FAUVEL

- Paratergites impunctate or finely and sparsely puntctate..................................................59

59 Antennal segment 10 about 3x as long as broad.: E (figs 205, 206).: S8 (fig. 212). 3.5- 4.2mm (FB 1.9-2.1mm). Brazil....................................................... brigita HROMÁDKA

- Antennal segment 10 about 2x as long as broad...............................................................60

60 Larger, FB 2.2-2.4mm.: E (figs 269, 300).: S8 (fig. 249). 4.3-5.0mm). Brazil.............. ................................................................................................................ josefa HROMÁDKA

- Smaller, FB  2,2mm .........................................................................................................61

61: Head broader.: E (figs 256, 264).: Unknown. 4.0-4.5mm (FB 2.1mm). Brazil....... .................................................................................................................... inopinus nov.sp.

-: Head less broad.: E (figs 255, 263).: Unknown. 4.0-4.8mm (FB 2.1-2.2mm). Brazil....................................................................................................... inamoenus nov.sp.

62 Smaller, FB 1.7-1.9mm.: E (fig. 169).: S8 (fig. 276). 3.5-4.2mm). Brazil................... ............................................................................................................ klimschi BERNHAUER

- Larger, FB 2.1-2.3mm.: E (figs 367, 368).: S8 (fig. 259). 4.0-5.0mm. Brazil, French Guiana, Suriname......................................................................... xenia HROMÁDKA

63 Frons broader (HW: DE <1.85).........................................................................................64

- Frons less broad (HE: DE 1.90)......................................................................................65

64 Complete frons very densely punctate (fig. 48).: E (fig. 170).: S8 (fig. 355). 3.8- 4.8mm (FB 2.2-2.3mm). Bolivia.............................................................. spissifrons PUTHZ

- Median portion of frons narrowly impunctate posteriorly (fig. 53).: E (figs 311, 318).: Unknown. 4.5-5.2mm (FB 2.4mm). French Guiana............. multidentatus nov.sp.

65 Head boader (HW: EW>1.02)..........................................................................................66

- Head less broad (HW:EW 0.97).....................................................................................67

66 Punctuation of T7 less coarse and less dense (fig. 96).: E (fig. 226).: S8 fig. 277). 3.8-4.5mm (FB 1.9-2.1mm). Panama, Suriname, BritishGuyana, Brazil, Ecuador............... ................................................................................................................ densipennis SHARP

- Punctuation of T8 coarser and denser (fig. 97).: E (fig. 306).: S8 about as in fig. 278. 3.8-4.5mm (FB 2.1-2.2mm). Costa Rica............................................. longinoi nov.sp.

67 Punctuation/sculpture of elytra irregularly rugose.: E (figs 309, 316).: S8 (fig.282). 3.7-4.5mm (FB 2.2-2.3mm). Ecuador, Peru........................... martae HROMÁDKA

- Punctuation/sculpture of elytra very dense, regular.: E (figs 112, 113, 330, 336).: S8 (fig. 340). 4.2-5.5mm (FB 2.3-2.4mm). Guatemala, El Salvador, Mexico, Panama........ .............................................................................................................. scabripennis SHARP

68 Whole abdomen reticulate.................................................................................................69

- Anterior tergites of abdomen not reticulate (tergite 7 reticulate or not reticulate.............87

69 Elytra reticulate.................................................................................................................70 Some species are listed twice since their reticulation may become indistinct

- Elytra not reticulate...........................................................................................................75

70 Elytral spot very large, broader than half the breadth of one elytron (figs 134, 150, 152)...................................................................................................................................71

- Elytral spot smaller, at most as broad as half the breadth of one elytron..........................74

71 Punctures of elytra discrete...............................................................................................73

- Punctures of elytra sometimes coalescent or connected by longitudinal striae (especially at the sides) (figs 77, 78).................................................................................72

72 Elytra (fig. 134).: E (fig. 188).: S8 (fig. 194). 5.9-6.5mm (FB 2.8-2.9mm). Peru........ ..................................................................................................... aurantiacoornatus PUTHZ

- Elytra (fig. 150).: E (fig. 161). 5.2-6.2mm (FB 2.9mm). Bolivia....................................... .................................................................................................... ( rufoornatus BERNHAUER )

73 spot often blurred.: E (fig. 271).: S8 as in fig. 247). 5.0-6.0mm (FB 3.0-3.2mm). Ecuador.................................................................................................. mariae HROMÁDKA

- Elytra (fig. x).: Elytral spot more distinct (fig. 152).: E (fig. x).: S8 (fig. 247). 6.0-6.5mm (FB 3.2mm). Ecuador............................................................... haravec nov.sp.

74: E (fig. 207).: S8 as in fig. 194. 5.0-6.2mm (FB 3.0-3.2mm). Venezuela, Ecuador, Bolivia.......................................................................................................... brooksi nov.sp.

-: E (fig. 304).: S8 (fig. 279). 5.0-6.0mm (FB 2.8-3.0mm). Ecuador.............................. ....................................................................................................... (lateralistriatus nov.sp.) 75 Elytral spot larger, broader than half of one elytron.........................................................76

- Elytral spot smaller, less broad than half of one elytron (fig. 126).: E (fig. 158).: S8 about as in fig. 282. 5.0-6.5mm (FB 2.5-2.7mm). Panama, Costa Rica electriger PUTHZ

76 Punctures on elytral spot vorticose forming a rosetta (fig. 79)..........................................77

- Punctures on elytral spot otherwise...................................................................................79

77 Elytral spot larger (figs 149, 151).....................................................................................78

- Elytral spot smaller (fig. 148).: Unknown.: S8 about as in fig. 359). 7.0-8.0mm (FB 3.8-4.0mm). Panama................................................ pulchriguttatus PUTHZ

78 Punctures of pronotum transversely confluent.: E (fig. 337).: Unknown. 7.0- 9.0mm (FB 3.8-4.0mm) Bolivia............................................................ pulchricolor PUTHZ

- Punctures of pronotum discrete.: E (fig. 160).: S8 about as in fig. 282). 7.5- 9.3mm (FB 4.1-4.3mm) Peru............................................................... pulchripennis PUTHZ

79 Elytra (fig. 133). Sternum 9 with a very long apicolateral tooth (about as in fig. 136).: E (fig. 166, 233).: S8 (fig. 231). 5.0-6.0mm (FB 2.6mm). Peru, Bolivia..................... ................................................................................................................. cuzcoensis PUTHZ

- 9 apicolaterally blunt or serrate or only with a short tooth................................................80

80 Elytral spot extending to the suture...................................................................................81

- Elytral spot not extending to the suture.............................................................................82

81 Elytra (fig. 122). Punctuation of pronotum very coarse and very dense, sometimes transversely coalescent.: (E (figs 175, 176, 178).: S8 (fig. 195). 5.2- 6.0mm (FB 2.8-2.9mm). Ecuador..................................................................................... alpaca nov.sp.

- Punctures of pronotum less coarse and less dense, discrete. E (fig. 304).: S8 (fig. 279). 5.0-6.0mm (FB 2.8-3.0mm). Ecuador...................................... lateralistriatus nov.sp.

82 Punctuation of pronotum dense, interstices at most as large as half diameter of punctures...........................................................................................................................83

- Punctuation of pronotum less dense, interstices mostly larger than half diameter of punctures...........................................................................................................................84

83 Elytra (fig. 66). Pronotum matt, punctuation extremely dense.: Unknown.: S8 about as in fig. 199. 7.0-8.2mm (FB 4.0mm). Peru........................ praecellens BERNHAUER

- Elytra (fig. 136). Pronotum shiny, punctuation less dense, interstices discrete.: E (fig. 167).: S8 about as in fig. 199. Peru, Bolivia, Argentina................. flavotaeniatus PUTHZ

84 Head broader (HW: EW <0.90)........................................................................................85

- less broad (HW: EW>0.90)...............................................................................................86

85: Elytral punctuation slightly coarser and denser (fig. 40, PUTHZ, 2005).: E (fig. 43, PUTHZ, l. c.; about as in fig. 242).: Unknown. 5.5-6.5mm (FB 3.2mm). Peru............ ............................................................................................................ flavofasciatus PUTHZ

- Elytral punctuation slightly less coarse, finer (fig. 39, l. c.).: E (fig. 242).: S8 about as in as fig. 194). 5.5-7.0mm (FB 3.1-3.2mm). Peru.................... flavocinctus PUTHZ

86: Elytral punctuation slightly coarser and denser, interstices less than twice as large as punctures.: E (fig. 237).: S8 (fig. 245). 5.0-6.0mm (FB 2.9-3.0mm). Bolivia, Peru................................................................................................. fossipennis BERNHAUER

- Elytral punctuation slightly less coarse, finer, interstices twice or more as large as punctures.: E (fig. 161). 5.2-6.2mm (FB 2.9mm). Bolivia.................. rufoornatus PUTHZ

87 Tergite 7 and 8 reticulate (can be very shallow on tergite 7)............................................88

- Whole abdomen without reticulation................................................................................97

88 Elytral spot larger, as broad or broader than half of one elytron.......................................89

- Elytral spot smaller, less broad than half of one elytron (fig. 137).: E (fig. 305).: S8 (fig. 278). 4.5-5.0mm (FB 2.4mm). Ecuador..........................................(llayca nov.sp.)

89 Elytral spot distinct...........................................................................................................90 - Elytral spot indistinct, becoming blurred in the ± dark brownish colour of elytra ...see # 44 ...............................................................................................................[ janae HROMÁDKA ]

90 Elytral spot extending to the side margin of elytra............................................................91

- Elytral spot not extending to the side margin of elytra.: Unknown.: S8 (fig. 297). 4.2-5.0mm (FB 2.6mm). Peru.......................................................................... palla nov.sp.

91 Punctuation of pronotum very dense and/or rugose, interstices much smaller than half diameter of punctures........................................................................................................93

- Punctuation of pronotum less dense, discrete, interstices less smaller than half diameter of punctures, shiny ............................................................................................................92

92 Elytral spot larger, extending to the suture (fig. 135). Larger: FB 2,9mm.: E (fig. 243). 6.0mm. Bolivia........................................................................ flavocingulatus PUTHZ

- Elytral spot smaller, not extending to the suture. Smaller, FB 2.6mm.: E (figs 182, 187).: S8 (fig. 197). 5.0-5.5mm (FB 2.6mm). Colombia......... aquatalensis BERNHAUER

93 Punctures on elytral spot coalescent (elytra: fig. 149).: E (figs 361, 375).: S8 (fig. 344). 5.0-6.0mm (FB 2.7-2.8mm). Ecuador............................................... sperlingi nov.sp.

- Punctures on elytral spot not coalescent............................................................................94

94 Smaller, FB <2,5mm. Elytra (fig. 145).: E (fig. 241, slightly tipped over). 4.5- 4.8mm (FB2.3mm). Colombia....................................................................... falcifer PUTHZ

- Larger, FB  2,9mm ...........................................................................................................95

95 Elytral spot smaller, oval (fig. 138).: Unknown.: S8 (fig. 294). 5.7-6.8mm (FB 3.1mm). Venezuela.............................................................................. obliquegutta nov.sp.

- Elytral spot larger, round, nearly as in fig 139..................................................................96

96: E as in S. amauta (figs 184, 185).: S8 about as in fig. 194. 5.5-6.5mm (FB 2.9- 3.0mm). Ecuador.............................................................................................. mauta PUTHZ

-: E (figs 184, 185).: S8 as in S. mauta . 5.5-6.5mm (FB 3.0-3.3mm). Ecuador.............. ...................................................................................................................... amauta PUTHZ

97 Sternum 9 with a very long tooth apicolaterally (as in fig.139)........................................98

- Sternum 9 apicolaterally with a short, ± blunt tooth and/or serrate...................................99

98 Pronotum and elytra with shallow reticulation. Larger, FB 3.0mm. E (fig. 338).: Unknown. 5.2-6.2mm. Colombia............................................................ sagittipenis PUTHZ

- Pronotum and elytra without reticulation. Elytra (figs 143, 146). Smaller, FB 2.5- 2.6mm.: E (figs 183, 332).: S8 (fig. 324). 4.0-5.0mm. Colombia, Ecuador.................. .......................................................................................................... postnotatus L. BENICK

99 Elytral spot broader than half the breadth of one elytron................................................100

- Elytral spot less broad than half the breadth of one elytron............................................102

100 Punctuation of elytral spot ± vorticose............................................................................101

- Punctuation of elytral spot discrete.: E (fig. 159).: S8 (fig. 193). 4.5-6.0mm (FB 2.6mm). Ecuador, Colombia............................................................................... apo PUTHZ

101 Larger, FB 3,7mm, sculpture of elytral spot rosetta-like (fig. 79).: S8 about as in fig.198 or 247). 6.5-7.5mm (FB 3.7-3.8mm). Colombia, Ecuador..... squamosus L. BENICK

- Smaller, FB 2.7mm, sculpture of elytral spot vorticose but not rosetta-like (fig. 77, 139).: E (figs 253, 261).: S8 (fig. 273). 4.6-5.6mm (FB 2.4-2.5mm). Venezuela......... ............................................................................................................. hermanianus nov.sp.

102 Elytral spot as in fig. 147, extending to the posterior margin of elytra.: E (fig. 163).: S8 (fig. 342). 4.3-5.6mm (FB 2.4-2.6mm). Venezuela........................ setipennis PUTHZ

- Elytral spot otherwise, not extending to the posterior margin of elytra..........................103

103 Abdominal punctuation denser, interstices as most as large as punctures.......................105

- Abdominal punctuation sparser, interstices distinctly larger than punctures...................107

104 Median portion of frons strongly elevated, smooth.........................................................104

- Median portion of frons slightly elevated, densely punctate...........................................106 105 Head broader (HW: EW 1.05), elytral spot larger, in lateral half of elytra (fig. 156).: E (fig. 339, 360).: S8 (fig. 343). 3.7-4.3mm (FB 2.1-2.2mm). Ecuador...................... ....................................................................................................................... simius nov.sp.

- Head less broad (HW: EW 0.94), elytral spot smaller, about in the middle of elytra (fig. 132).: E (fig. 227). 4.0-4.5mm (FB 2.2mm). Costa Rica, Panama............................. ............................................................................................................ ( cruentus L. BENICK )

106 Smaller, FB 2.0-2.3mm, head broader (HW: EW 1.13). Elytra (fig. 154).: E (fig. 172).: S8 (fig. 293). 3.5-4.5mm. Peru........................................................... nusta PUTHZ

- Larger. FB 2.2-2.4mm, head less broad (HW: EW 1.06).: E (figs 217, 225).: Unknown. 4.0-4.7mm (FB 2.4mm). Ecuador.................................................. cholo nov.sp.

107 Larger, FB 2.4mm.........................................................................................................108

- Smaller, FB  2,3mm .......................................................................................................113

108 Elytral spot larger (figs 131, 155)...................................................................................109

- Elytral spot smaller (figs x, y).........................................................................................110

109 Head broader (HW: EW>1.00), elytra (fig. 131).: E (fig. x). 4.2-4.5mm (FB 2.4- 2.5mm). Bolivia........................................................................... cochabambaensis nov.sp.

- Head less broad (HW: EW 0.96), elytra (fig. 155).: E (fig. 327).: S8 (fig. 296). 4.3-5.0mm U(FB 2.4-2.5mm). Colombia................................................. ovatigutta nov.sp.

110 Paratergites 5 densely punctate, elytral spot nearly indistinct.: E (figs. 312, 319).: S8 about as in fig. 196). 4.5-5.5mm (FB 2.4-2.5mm). French Guiana.................................. ........................................................................................................ obscuroguttatus nov.sp.

- Paratergites 5 impunctate or only with few punctures....................................................111

111 Elytral spot in lateral half of elytra (figs 137, 141).........................................................112

- Elytral spot in about the middle of elytra (fig. 142).: E (figs 329, 334).: S8 (fig.322). 4.0-5.0mm (FB 2.2-2.3mm). Costa Rica............................... parvuligutta nov.sp.

112 Lateral portions of frons coarsely and very densely punctate; pronotum less uneven, elytra (fig. 141).: E (fig. 328).: S8 (fig. 298). 4.7-5.6mm (FB 2.5- 2.6mm). Ecuador.................................................................................. ovatiguttatus nov.sp.

- Lateral portions of frons moderately finely and moderately densely punctate, pronotum conspicuously uneven, elytra (fig. 137) .......................................................(llayca nov.sp.)

113 Elytral spot larger............................................................................................................114

- Elytral spot smaller (fig. 132).: E (fig. 227). 4.0-4.5mm (FB 2.2mm). Costa Rica, Panama................................................................................................... cruentus L. BENICK

114 Head broader (HW: EW>1.00)......................................................................................115

- Head less broad (HW: EW <0.97)...................................................................................116

115 Elytral punctuation coarser, diameter of punctures larger than cross section of antennal segment 2. Punctuation of the lateral portions of frons very coarse, diameter of punctures about as large as apical cross section of antennal segment 2.: E (fig. 310, 317).: Unknown. 3.5-3.7mm (FB 2.1mm). Peru.................................... mazureki nov.sp.

- Elytral punctuation less coarse, diameter of punctures about as large as apical cross section of antennal segment 2. Elytra (fig. 127). Punctuation of the lateral portions of frons less coarse, diameter of punctures smaller than diameter of antennal segment 3.: E (fig. 201).: S8 (fig. 198). 3.4-3.7mm (FB 1.8-2.0mm). Venezuela.......................... ..................................................................................................................... bolivari nov.sp.

116 Paratergites 5 densely punctate.......................................................................................117

- Paratergites 5 sparsely punctate......................................................................................118

117: E. Antennal segment 10 1/3 longer than broad. Elytra (fig. 140).: E (figs 313, 321).: S8 as in fig. 198. 3.5-4.0mm (FB 2.0-2.1mm). Venezuela.............. ojedai nov.sp.

-: E. Antennal segment 10 1.5x longer than broad.: E (fig. 363, 373).: Unknown. 4.2mm (FB 2.3mm). Venezuela................................................................ vespuccii nov.sp.

118 Elytral spot larger (fig. 144), elytral punctuation denser, confluent in posterior half.: E (fig. 260). 4.3-5.0mm (FB 2.2-2.3mm). Colombia...................................... gibbus PUTHZ - Elytral spot (fig. 129), elytral punctuation less dense, less confluent in posterior half.: E (fig. 202, 209).: S8 as in fig. 198. 4.0-4.7mm (FB 2.2- 2.3mm). Ecuador............... ............................................................................................................... cavatigutta nov.sp.