Calvarium (Calvariellum)
|
publication ID |
https://doi.org/ 10.11646/zootaxa.3846.1.1 |
|
publication LSID |
lsid:zoobank.org:pub:97D4A04A-D75E-45CC-8A70-3EB3A4E94D9B |
|
DOI |
https://doi.org/10.5281/zenodo.6126807 |
|
persistent identifier |
https://treatment.plazi.org/id/113287AD-9B5F-FFDB-FF13-FC00FCA6FC1F |
|
treatment provided by |
Plazi |
|
scientific name |
Calvarium (Calvariellum) |
| status |
|
Calvarium (Calvariellum) , new subgenus
Type species: Calvarium (Calvariellum) hamifer , n. sp.
Diagnosis. Small (BL 1.7 − 2.2 mm), parallel-sided beetles exhibiting the before-mentioned generic characters but distinguished by the following:
• Head without subocular ridge, about 1.7 times wider across eyes than long from occipital foramen to anterior edge of frontoclypeus ( Fig. 6 View FIGURES 1 − 9 ).
• A transverse arched suture subdividing the gula ( Fig. 6 View FIGURES 1 − 9 ).
• Penis with wide triangularly excised pala, large trigonium, and simple parameroids ( Figs. 24 View FIGURES 21 − 25 , 42 View FIGURES 38 − 42 ), or parameroid tips caudally connected ( Figs. 30, 36 View FIGURES 29 − 37 ).
• Tegmen and parameres Y-shaped (e.g., Fig. 25 View FIGURES 21 − 25 ), or parameres separate, no visible tegmen (e.g., Fig. 31 View FIGURES 29 − 37 ). • T9 not sclerotized, only the long rod-like apodemes distinct. They may be connected by a membrane ( Fig. 34 View FIGURES 29 − 37 ), or completely separate (e.g., Fig. 23 View FIGURES 21 − 25 ).
• T8 of two species with paired ventral appendages ( Figs. 21 View FIGURES 21 − 25 , 29 View FIGURES 29 − 37 ).
• S8 indistinct or very weak, S9 lightly sclerotized.
Description. Elongate, sides of elytra parallel. In dorsal view, almost no angle between elytra and pronotum, none between pronotum and head ( Fig. 18 View FIGURES 18 − 20 ). Head very wide, distance between eyes large. In frontal view, an eye, the labrum and the 1st antennal segment each occupy roughly 1/5 of total head width ( Figs. 6, 7 View FIGURES 1 − 9 ).
Antennal sulcus between eye and mouthparts deep, bordered by high crests. The medial crest runs from the inner edge of the antennal sulcus up to above the mandibular articulation. The frontal ridge above the scape turns into the outer crest which descends close to the eye. The outer crest eventually meets the subgenal ridge at the point where a short transverse branch connects it to an arched accessory suture ( Fig. 7 View FIGURES 1 − 9 ).
Antennal base as for genus. Scape a little longer than pedicel. Pedicel and first flagellar segment of similar length, segment 4 thin, cylindrical. Flagellum loosely structured, segments constricted at both ends. Last segment elongate oval ( Fig. 19 View FIGURES 18 − 20 ).
Mandibles symmetrical, with distinct tooth. Last segment of maxillary palpus short, bottle-shaped. Second segment of labial palpus curved, its end with the concave terminal segment directed mediad ( Figs. 6, 8, 9 View FIGURES 1 − 9 ). Prosternal process anteriorly parallel, caudally widened, rear edge regularly rounded ( Fig. 20 View FIGURES 18 − 20 ), sclerite punctate and finely pilose.
Receiving groove on mesoventrite wide, almost hexagonal, bordered by low ridges. Mesoventral process wide and short, not extending to rear edge of middle coxae which are widely separated by a finely bordered parabolic process of the metaventrite ( Fig. 20 View FIGURES 18 − 20 ). Ventral face of thorax with coarse setiferous punctures except in places where the femora rest. Discrimen of metaventrite complete.
Legs unmodified. Abdominal sternites unmodified. A few setiferous granules on the small triangle formed by S3 between the hind coxae, a long granule row along the front edge of S4.
Male. T8 with only microtrichia, no setal fringe (e.g., Figs. 29 View FIGURES 29 − 37 , 39 View FIGURES 38 − 42 ). S8 weakly developed, V-shaped ( Figs. 29, 33 View FIGURES 29 − 37 ). Plate of T9 not developed, apodemes caudally prolonged, connected ( Fig. 34 View FIGURES 29 − 37 ) or separate ( Figs. 23 View FIGURES 21 − 25 , 40 View FIGURES 38 − 42 ). S9 with a slender base and an oval, caudally sparsely pilose bilobed or rounded plate (e.g., Figs. 23 View FIGURES 21 − 25 , 35 View FIGURES 29 − 37 ). Penis stout, pala anteriorly wide, excised, with anterolateral corners (e.g., Figs. 24 View FIGURES 21 − 25 , 36 View FIGURES 29 − 37 ). Trigonium and parameroids well developed, variable between species. The parameres are slender blades attached to a narrow handle-like tegmen (e.g., Figs. 25 View FIGURES 21 − 25 , 37 View FIGURES 29 − 37 ).
Female. No hair tuft or other modification to abdominal sternites. T8 with long and strong apodemes, those of the hairless small S8 much shorter, thin, anteriorly connected by a loop. Ovipositor unmodified ( Fig. 26 View FIGURES 26 − 28 ), the long apodemes with short recurrent branchlet. Prehensor variably developed ( Figs. 27, 28 View FIGURES 26 − 28 ). No distinct pores on bursella or gonoduct.
Note. All four species agree with the foregoing description which is not repeated under individual species. Only short diagnoses of distinctive traits are provided, in order of descending similarity with the type species. By their genitalia, the species are recognized at a glance, no key is needed.
Etymology. Calvariellum, diminutive of Calvarium .
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.