Calvarium (Calvariellum) cochlearifer, Zwick, Peter, 2014

Zwick, Peter, 2014, Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer, Zootaxa 3846 (1), pp. 1-41 : 11-12

Calvarium (Calvariellum) cochlearifer , n. sp.

( Figs. 28–31 )

Type material. Holotype: 1♂: “ 15.03S 145.09E 3km NE of Mt. QLD Webb 2 Oct. 1980 D.H.Colless (Malaise Trap)” ( ANIC). Paratypes (all ANIC): 1♂: “ 15.03S 145.09E 3km NE of Mt. QLD Webb 30 Apr. 1981 D.H.Colless (at light)”; 1♂: “Moses Ck 4 km NbyE of Mt Finnigan QLD 15 Oct. 1980 D.H.Colless”; 1♂: “ 15.30S 145.16E 1 k SE of Mt. Cook QLD 13 Oct.1989 D.H.Colless”; 2♂: “ 15.03S 145.09E 3km NE of Mt. Webb QLD 30 Apr. 1981 D.H.Colless (at light)”; 1♂ 1♀: “Bamboo Ck., near Miallo [16° 23.392' S 145° 22.503' E] N. of Mossman N.Qld. 25 Apr. 1967 D.H. Colless”.

Habitus. Similar to C. hamifer . BL 1.9–2.2 mm. Dark brown, appendages lighter.

Male. T8 only with microtrichia, caudal edge slightly angular. Disc with grouped microtrichia, very short setae only near the caudolateral angles. Apodemes less than twice as long as plate, front edge of plate barely defined. Ventrally near the end of each apodeme is an irregularly shaped sclerotized pocket. S8 with wide U-shaped base, disc unsclerotized, bare ( Fig. 29 ).

T9 represented only by long rod-shaped apodemes ending in sharp tips. S9 with paired sclerotized strips in the narrow base, oval plate caudally vaguely bilobed, with lateral groups of fine setae. Tegmen with unpaired slender base, parameres in the form of slender gently inwardly curved barely sclerotized horns ( Fig. 31 ). Penis base squarish, front slightly concave, lateral corners blunt. Trigonium stout, a little inflated near midlength, ending in short lateral points and a median spoon-shaped appendage with downcurved blunt tip. The parameroids are longer than the trigonium. Their slender bases curve away from the trigonium but their wide pore-bearing flat tips overlap dorsally from the trigonium ( Fig. 30 ).

Female. A flat structure in the gonoduct consisting of two lateral sclerites with wide-lobed overlapping tips seems to be the prehensor ( Fig. 28 ).

Etymology. The Latin noun in apposition translates as spoon-carrier (cochlear = spoon, Latin), a reference to the shape of the penis apex.

Note. Known only from tropical North Queensland. The outgrowth from the ventral side of T8 and the strongly curved appendage at the penis tip suggest C. hamifer and C. cochlearifer are sister taxa. They were taken together at Bamboo Creek, the association of sexes based on common occurrence seems nevertheless reliable.

Figures

FIGURES 26 − 28. Calvarium (Calvariellum) spp., females: 26, C. hamifer, n. sp. (paratype from Station Ck), abdomen with protracted ovipositor and prehensor in situ; 27, prehensor enlarged, diagrammatic. 28, C. cochlearifer, n. sp. (from Bamboo Ck), prehensor. Abbreviation: cf, circular fold. 27 and 28 to the same scale.

FIGURES 29 − 37. Calvarium (Calvariellum) spp., males. C. cochlearifer, n. sp.: 29, T 8 and S 8, superimposed; 30, penis; 31, T 9, S 9 and tegmen (tn) with parameres (p) superimposed. C. bellendenker, n. sp.: 32, T 8; 33, S 8; 34, T 9; 35, S 9; 36, penis; 37, tegmen and parameres. Figures for a given species to the same scale.