Ypsiloncyphon katherinae, Zwick, Peter, 2014

Zwick, Peter, 2014, Australian Marsh Beetles (Coleoptera: Scirtidae). 6. Genera Calvarium Pic, Papuacyphon Zwick, and Ypsiloncyphon Klausnitzer, Zootaxa 3846 (1), pp. 1-41 : 31-32

Ypsiloncyphon katherinae , n. sp.

( Figs. 103–109 )

Type material. Holotype ♂: “Katherine Gorge NT 26.x.75 light trap M.J.Muller”. Paratypes, NT: 1♂: “Caranbirini Waterhole, 16.16S 136.05E, 33 km SW of Borroloola, N.T., 21.iv.1976, at light, J.E.Feehan”. 1♂: “ 16.02S 130.27S NT Gregory N.P. 8.3km N Humbert Junction 5–12 June 2001 M.Irwin, F.Parker, C.Lambkin \ Malaise in dry bed near flowing creek“. QLD: 1♂: “ 18.36S 138.08E QLD Musselbrook Camp 8–21 May 1995 I.D.Naumann at light“. 1♂: “ 18.33S 138.11 E GFS Holts Ck 8km N of Musselbrook Mining Camp QLD 10–20 May 1995 T.Weir at light“. 1♂: “ 18.37S 137.59 GPS Border Waterhole, Musselbrook 15km W by S Musselbrook Mining Camp ( QLD) NT 14 May 1995 T.Weir at light“. (all ANIC).

Additional material studied. NT: 1♀(presumed): “ 16.08S 136.06E 22 km WSW of Bonoloola, NT, 2 Nov.1975 M.S.Upton”. 1♀(presumed): “ 16°8'52''S 130°26'38''E GPS NT Gregory NP Humbert Track 4.4km SSE Bullita Camp Ground 12-14 Jun 2001 ME.Irwin, RD.Parker, C.Lambkin \ Malaise trap 17b white sand” (all ANIC). 1♀(presumed): “ Australia: N.T. 100 km N of Tennant Ck 8.v.1987 Fay & Halfpapp at light” (QDPC-M).

Habitus. An elongate oval species, not all males stouter than females. Male BL 1.9–2.1 mm, BL /BW varies from 1.7 to 2.1, female BL is 2.0– 2.2 mm, BL /BW ~2.

Male. Apodemes of T9 about 1.8 times longer than those of T8 (not shown). S9 with long, narrow capulus with triangular base. Half of the longer blade is wide, the other half a slender flagellum. The shorter blade is only little shorter, its wide part is longer, its terminal flagellum occupies only about 1/3 of the length ( Fig. 103 ). Tegmen and parameres divided down to the barely enlarged base ( Fig. 105 ). Penis ( Fig. 104 ): the stalk-like pala with large triangular base occupies about 40% of total length. Penis widening much where it divides into parameroids and trigonium, the latter is more than half as wide as total width. Trigonium strongly waisted near midlength, then again much widened to a short wide cap with terminal nipple. Curvature of the rod-like parameroids follows the contour of the trigonium: they converge, then diverge again, and finally curve over the trigonium apex.

Female (presumed). T7 with 2 paramedian notches of the same size and shape as the small nipple between them ( Fig. 107 ). Apodemes of S8 long and slender, only a short anterior portion fused, rod-like ( Fig. 106 ). Vulvar sclerite fairly large, pocket- or box-like, sides almost meeting ventrally, caudal portion with vaguely delimited sclerotized areas, details not visible in preparations ( Fig. 108 ). Dictyon of normal size but with only few meshes ( Fig. 109 ), the ones in the centre smaller than the surrounding ones, the mesh with the peripheral outgrowths is very large.

Notes and etymology. In Northwest Australia occur several species with long slender pala and capulus. Ypsiloncyphon katherinae is distinguished by sinuous rod-shaped parameroids, the important basal width of the penis, and the apically strongly enlarged trigonium with nipple-shaped apex. The notches on female T7 are unique. Sexes are tentatively associated because they occur in the same general area in the Northern Territorry, between the borders of Western Australia and Queensland, with both sexes taken in Gregory N.P. The name is a noun in the genitive case honouring the presumed name patron of the type locality.

Figures

FIGURES 103 – 109. Ypsiloncyphon katherinae, n. sp., male (103 – 105) and female (106 – 109): 103, ♂ holotype, S 9; 104, penis, dorsal; 105, tegmen and parameres; 106, ♀ (N of Tennant Ck.) terminal segments, apodemes of S 8 black; 107, ♀ T 7; 108, vulvar sclerite; 109, dictyon. Figs. 103 – 106 to the same scale.