Cletocamptus Schmankevitsch, 1875

Gomez, Samuel & Yanez-Rivera, Beatriz, 2022, The genus Cletocamptus (Harpacticoida, Canthocamptidae): a reappraisal, with proposal of a new subfamily, a new genus, and a new species, ZooKeys 1080, pp. 165-208 : 165

Genus Cletocamptus Schmankevitsch, 1875

Type species.

Cletocamptus retrogressus Schmankevitsch, 1875 (type by original designation).

Other species.

Cletocamptus affinis Kiefer, 1957, C. albuquerquensis (Herrick, 1894), C. assimilis Gomez & Gee, 2009, C. axi Mielke, 2000, C. cecsurirensis Gómez, Scheihing & Labarca, 2007, C. chappuisi Gómez, Gerber & Fuentes-Reinés, 2017, Cletocamptus confluens (Schmeil, 1894), C. deborahdexterae Gómez, Fleeger, Rocha-Olivares & Foltz, 2004, C. dominicanus Kiefer, 1934, C. feei (Shen, 1956), C. fourchensis Gómez, Fleeger, Rocha-Olivares & Foltz, 2004, C. goenchim Gómez, Ingole, Sawant & Singh, 2013, C. gomezi Suárez-Morales, Barrera-Moreno & Ciros-Pérez, 2013, C. gravihiatus (Shen & Sung, 1963), C. koreanus Chang, 2013, C. levis Gómez, 2005, C. mongolicus Stĕrba, 1968, C. nudus Gómez, 2005, C. pilosus Gomez & Gee, 2009, C. samariensis Fuentes-Reinés, Zoppi de Roa & Torres, 2015, C. schmidti Mielke, 2000, C. sinaloensis Gómez, Fleeger, Rocha-Olivares & Foltz, 2004, C. spinulosus Gomez & Gee, 2009, C. stimpsoni Gómez, Fleeger, Rocha-Olivares & Foltz, 2004, C. tainoi Gómez, Gerber & Fuentes-Reinés, 2017, C. tertius Gomez & Gee, 2009, C. trichotus Kiefer, 1929.

Species incertae sedis.

Marshia brevicaudata Herrick, 1894.

Species inquirendae.

Cletocamptus bermudae Willey, 1930, C. cfr. bicolor sensu Herbst (1960), C. brehmi Kiefer, 1933, Cletocamptus deitersi (Richard, 1897), C. deitersi sensu Chappuis (1934), C. deitersi sensu Daday (1902), C. deitersi sensu Dussart (1974), C. deitersi sensu Hamond (1973), C. deitersi sensu Herbst (1960), C. deitersi sensu Kiefer (1936), C. deitersi sensu Suárez-Morales et al. (1996), C. deitersi sensu Tai and Song (1979), C. ecuadorianus Löffler, 1963, C. gabrieli Löffler, 1961, C. kummleri (Delachaux, 1917), Godotella dadayi Delachaux, 1917.

Doubtful records.

C. deitersi : in Apostolov (1984), Brehm (1936, 1965), Chappuis (1936), Dussart and Frutos (1986), Loftus and Reid (2000), Oliveira et al. (1971), Ranga Reddy and Radhakrishna (1979), Ringuelet (1958a, 1958b, 1960, 1962), Ringuelet et al. (1967), Ruber et al. (1994), Sitjar (1988), Zamudio-Valdéz (1991).

Diagnosis.

Canthocamptidae : Cletocamptinae . Body fusiform, without clear distinction between prosome and urosome. Without nuchal organs on cephalothorax or body somites. Female rostrum distinct, rarely fused to cephalothorax, large and triangular with broad proximal margin; ornamented with ventral (sub)distal spinules and with rounded tip in both sexes. Posterior margin of cephalothorax and/or prosomites with long and slender, or short spinules, or serrated; posterior margin of urosomites except for anal somite with short spinules or serrated; body somites without cuticular sensillum-bearing socles; anal operculum without dorsal ornamentation or with transverse row of strong, large, or short, small spinules; posterior margin unornamented, or with small or large spinules. Female genital somite and third urosomite separated dorsolaterally, completely fused ventrally forming genital double-somite. Female antennule six-, rarely seven-segmented (the latter reported for C. gravihiatus requires confirmation). Antenna with allobasis, with one or two abexopodal setae (proximal element basal, distal seta endopodal); exopod one-segmented and longer than wide, or minute, rarely represented by single seta (the latter reported for C. chappuisi requires confirmation). Mandibular palp one-segmented, rarely two-segmented (basis and endopod distinct as in C. retrogressus ); when one-segmented, then very small and wider than long, or as long as wide ( C. dominicanus ), or longer than wide ( C. confluens , C. gomezi ); when palp one-segmented, then basis and endopod not discernible, rarely discernible (represented by single seta as in C. confluens and C. dominicanus ); endopod with two setae, rarely with one single element ( C. pilosus ); exopod absent; with or without surface (most probably exopodal) seta on coxa, the latter present only in some species with a one-segmented palp wider than long, absent in species with palp as long as wide or longer than wide. Maxillule with endopod and exopod incorporated to basis; praecoxal arthrite with ventral seta thick and strongly spinulose, or slender and pinnate or smooth. Maxilla with two syncoxal endites; endopod completely incorporated to allobasis. Maxilliped subchelate; syncoxa with one seta; basis unarmed; claw with accessory seta. P1ENP not prehensile; P1-P4EXP three-segmented; female P1-P3ENP two-segmented, P4ENP two- or one-segmented; inner exopodal and endopodal setae with or without comb tip (see Gómez et al. 2017). Female P5EXP and BENP fused; both baseoendopods of P5 separated. Armature formulae as follows:

Female P6 with one or two setae. Caudal rami with six or seven setae; setae IV and V fused basally or separated.

Sexual dimorphism expressed in a) rostrum (slenderer than in female), b) the male antennule (subchirocer), c) basis of P1 (with inner distal process), d) outer spine on P2 ENP2 (thicker and/or shorter than in the female), e) P3ENP (three-segmented as in most species, or two-segmented as in C. albuquerquensis , C. chappuisi , C. confluens , C. dominicanus , and C. tainoi ; when three-segmented then inner apophysis on second segment; when two-segmented then inner apophysis medially as in C. dominicanus , or subdistally on second segment as in C. albuquerquensis , C. chappuisi , C. confluens , and C. tainoi , but occasionally with additional outer apophysis as in C. confluens ), f) P5 (both legs fused medially as in most species, or separated as in C. axi , C. cecsurirensis , C. retrogressus , and C. schmidti ; EXP and BENP fused, and both legs articulating with somite, g) P6 (composed of two lappets articulated to somite and unarmed as in most species or, occasionally, with one seta attributable to intraspecific variability as in C. axi , C. schmidti , C. sinaloensis ). Additionally, sexual dimorphism can be expressed also (depending on the species) in a) P2EXP and/or P3 and P4EXP (segments longer than in the female, outer spines stronger than in the female, and/or rami curved inwards as in C. retrogressus , C. confluens , C. albuquerquensis , C. samariensis , C. spinulosus , C. tainoi , C. trichotus ), b) shape of P2-P4ENP (segments thicker than in female as in C. confluens ), c) caudal rami (longer than in the female).