Charpentieria itala allatollae ( Kaeufel , 1928)

De Mattia, Willy, Reier, Susanne & Haring, Elisabeth, 2021, Morphological investigation of genital organs and first insights into the phylogeny of the genus Siciliaria Vest, 1867 as a basis for a taxonomic revision (Mollusca, Gastropoda, Clausiliidae), ZooKeys 1077, pp. 1-175 : 1

Charpentieria itala allatollae ( Kaeufel, 1928)

Figs 44.7, 44.8, 47.6

Distribution.

This taxon is known to occur in Val Ampola and Val Lorina (Province of Trento, Trentino-Alto Adige, Italy). It is also reported by Gredler (1891) for Monte Tombea as Clausilia tombeana Gredler, 1891 and from other localities in eastern Lombardia by Nardi (2011: 101).

Specimens examined.

Italy, Trentino-Alto Adige, Trento, Storo, Albergo alla Tolla , Val Ampola. 720 m asl, 45°51'7.62"N, 10°38'4.51"E, W. De Mattia and J. Macor leg. and det., 2 dissected spm .

External morphology of the genital organs

(Fig. 44 .7). The FO is 1.5 × longer than the V. The FDBC is longer than the BC+SDBC. The BC+SDBC is club-like, with visible distinction between the SDBC and the BC. The apex is rounded. The D is slightly longer than the BC+SDBC, thin and with a round apex. The V is short but wide in diameter. The PC is double as long as the V. The P is swollen along its whole length. The PR is short and robust. The E is longer than the P but thinner in diameter.

Internal morphology of the genital organs

(Fig. 44 .8). The A, the V and the P are smooth, with few, very loose and barely visible longitudinal elevations. The PP is almost smooth, elongated with a pointed apex. The PP originates from the distal end of the ELP. The ER is not present. The epiphallar formula is: PP(ELP). The E shows two ELP that proximally fade before the VD.

Remarks.

This taxon, previously considered as a junior synonym of C. itala lorinae ( Nordsieck 1963b: 184) was recently elevated (with no explanation) as a valid subspecies by the same author ( Nordsieck 2007: 54). It was long considered as a transitional hybrid form between C. clavata and C. itala but this assumption was based exclusively upon shell characters ( Nardi 2011: 101). Scheel and Hausdorf (2012: 3799) proved this is not correct as the taxa previously belonging to one or the other species appear deeply intermingled in the phylograms they provided.

Figures

Figure 44. Charpentieria itala itala (G. von Martens, 1824), Barbarano Vicentino 44.1, 44.2 whole distal genital organs 44.3 internal distal part of genital organs 44.4 detail of the pseudopapilla. Charpentieria itala albopustolata (De Cristofori & Jan, 1832), Brescia 44.5 whole distal genital organs 44.6 internal distal part of genital organs. Charpentieria itala allatollae (Kaeufel, 1928), Val Ampola, Storo 44.7 whole distal genital organs 44.8 internal distal part of genital organs.