Allmonia funiculata ( Borson, 1821 ) Harzhauser & Landau, 2019

Harzhauser, Mathias & Landau, Bernard, 2019, Turritellidae (Gastropoda) of the Miocene Paratethys Sea with considerations about turritellid genera, Zootaxa 4681 (1), pp. 1-136 : 98-99

Allmonia funiculata ( Borson, 1821) new comb.

Figs 25 I–J

Turritella fasciata nob. — Borson, 1821: 343, pl. 2, fig. 12 [non Turritella fasciata Lamarck, 1804 ].

* Turritella funiculata nob. — Borson, 1821: 343, pl. 2, fig. 13.

Turritella cathedralis A. Br. — Brongniart, 1823: 55 (pars) [Italian specimens only; non pl. 4, fig. 6].

[ Turritella ] funiculata Borson — Krüger, 1825: 392.

Turritella (Proto) cathedralis Brong. var. trisulca Myl. — Mylius, 1891: 336, fig. 14.

P [rotoma]. cathedralis (Brong.) — Sacco, 1895: 31, pl. 3, fig. 12 [non Allmonia cathedralis ( Brongniart, 1823) ].

P [rotoma]. cathedralis var. pseudolaevis Sacc. — Sacco, 1895: 32, pl. 3, fig. 13.

P [rotoma]. cathedralis var. pseudovermicularis Sacc. — Sacco, 1895: 32, pl. 3, fig. 14.

P [rotoma]. cathedralis var. semilaevis Sacc. — Sacco, 1895: 32, pl. 3, fig. 15.

P [rotoma]. cathedralis var. unisulcata Sacc. — Sacco, 1895: 32, pl. 3, fig. 16.

P [rotoma]. cathedralis var. funiculata (Bors.) — Sacco, 1895: 32, pl. 3, figs 17a–17b.

P [rotoma]. cathedralis var. concavosimplex Sacc. — Sacco, 1895: 33, pl. 3, fig. 18.

P [rotoma]. cathedralis var. inflatosimplex Sacc. — Sacco, 1895: 33, pl. 3, fig. 19.

P [rotoma]. cathedralis var. exfasciata Sacc. — Sacco, 1895: 33, pl. 3, fig. 20.

P [rotoma]. cathedralis var. suprainflata Sacc. — Sacco, 1895: 33, pl. 3, fig. 21.

Turritella cathedralis A. Brongniart, 1823 — Cossmann, 1910b: 187, figs C–Ca.

T [urritella]. funiculata Borson — Cossmann, 1912: 129.

P [rotoma]. cathedralis var. pseudolaevis Sacc. — Sacco, 1904: 125, pl. 25, fig. 31.

P [rotoma]. cathedralis var. pseudovermicularis Sacc. — Sacco, 1904: 125, pl. 25, fig. 32.

? Protoma rotifera (Lamk.) — Stefanini, 1916: 95, pl. 2, fig. 16 (only) [non Allmonia rotifera ( Lamarck, 1822) ].

T [urritella]. funiculata Borson — Cossmann & Peyrot, 1922: 229.

Turritella fasciata— Pavia, 1976: 148.

Turritella funiculata— Pavia, 1976: 148.

Protoma cathedralis ( Brongniart, 1823) — Ferrero Mortara et al., 1984: 236 [non Allmonia cathedralis ( Brongniart, 1823) ].

[ Protoma cathedralis ( Brongniart, 1823) ] var. pseudolaevis Sacco, 1895 — Ferrero Mortara et al., 1984: 237.

[ Protoma cathedralis ( Brongniart, 1823) ] var. pseudovermicularis Sacco, 1895 — Ferrero Mortara et al., 1984: 237.

[ Protoma cathedralis ( Brongniart, 1823) ] var. semilaevis Sacco, 1895 — Ferrero Mortara et al., 1984: 237.

[ Protoma cathedralis ( Brongniart, 1823) ] var. unisulcata Sacco, 1895 — Ferrero Mortara et al., 1984: 237.

[ Protoma cathedralis ( Brongniart, 1823) ] var. funiculata ( Borson, 1821) — Ferrero Mortara et al., 1984: 237.

[ Protoma cathedralis ( Brongniart, 1823) ] var. concavosimplex Sacco, 1895 — Ferrero Mortara et al., 1984: 237.

[ Protoma cathedralis ( Brongniart, 1823) ] var. inflatosimplex Sacco, 1895 — Ferrero Mortara et al., 1984: 237.

[ Protoma cathedralis ( Brongniart, 1823) ] var. exfasciata Sacco, 1895 — Ferrero Mortara et al., 1984: 237.

[ Protoma cathedralis ( Brongniart, 1823) ] var. suprainflata Sacco, 1895 — Ferrero Mortara et al., 1984: 237.

[ Protoma cathedralis ( Brongniart, 1823) ] var. quadriplicata ( Basterot, 1825) — Ferrero Mortara et al., 1984: 237.

Protoma cathedralis (Brongniart, 1856) [sic] — Zunino & Pavia, 2009: 356, pl. 1, fig. 3 [non Allmonia cathedralis ( Brongniart, 1823) ].

Type material. According to Pavia (1976), the type specimens of Turritella fasciata Borson, 1821 and Turritella funiculata Borson, 1821 are lost. The plate caption for the specimen illustrated by Sacco (1996, pl. 3, fig 17a) as Protoma cathedralis var. funiculata , however, states: ‘es. tip. Coll. Borson’. Therefore, the specimen BS.061.07.006, stored in the collection of the Museo Regionale di Scienze Naturali Torino, was part of Borson’s collection and most probably represents a syntype or even the holotype. Merlino & Campanino (2002) list this specimen as syntype.

Studied material. 2 spec. NHMW 1862/0049/0026, 3 spec. NHMW 1854/0016/0209, 7 spec. NHMW 1862/0035/0454, Torino ( Italy) (the preservation suggests Baldissero as locality) .

Illustrated material. Fig. 25I : NHMW 1862/0049/0026, SL: 94.3 mm, MD: 20.4 mm; Fig. 25J : NHMW 1854/0016/0209 SL: 80.9 mm, MD: 26.3 mm.

Discussion. The first description of Italian specimens that we consider as representatives of Allmonia was published by Borson (1821) as Turritella fasciata and T. funiculata from the Burdigalian of the Torino Hills. Turritella fasciata was already preoccupied for an Eocene species described by Lamarck (1804) but Turritella funiculata was an available name. Borson (1821) provided a short description and a rather sketchy illustration. Brongniart (1823) was aware of this name but rejected its use by referring to the poor illustration in Borson (1821) and synonymized these with his Turritella cathedralis . Sacco (1895) restored Borson’s taxa as variation-names (≈ subspecies) of Protoma cathedralis as exfasciata (replacement name for fasciata ) and funiculata . Cossmann (1912) and Cossmann & Peyrot (1922) argued that the Italian Miocene specimens were different from the French cathedralis and pointed out that funiculata was an available name for the Italian species. Finally, Pavia (1976) discussed funiculata as an available name to replace some of Sacco’s variety names. In respect to this literature record, Turritella funiculata cannot be treated as nomen oblitum. Sacco (1895) described numerous additional variation names of P. cathedralis , which all derive from the Burdigalian locality Baldissero in the Torino Hills ( Italy). Therefore, we tentatively treat all these taxa as morphotypes of a single, very variable species for which Turritella funiculata is the oldest available name.

Distribution. Proto-Mediterranean Sea. Burdigalian (early Miocene): Torino Hills: Baldissero ( Italy). A fragmentary specimen illustrated by Stefanini (1916) from the Tortonian of the Veneto region in northern Italy would be the stratigraphically youngest record.

Figures

FIGURE 25. A–D. Allmonia paucicincta (Sacco, 1895), Burgschleinitz (Austria). A. Holotype, NHMW 1846/0010/0014. B1–B3. Collection Anton Breitenberger. C. NHMW 2018/0128/0002. D. NHMW 2018/0128/000. E. Turritella (Protoma) telleri Hilber, 1993, holotype, Ivanji dol (Slovenia), UMJG&P 1247. F–H. Allmonia cathedralis (Brongniart, 1823), Léognan (France). F1–F2. NHMW 1858/0018/0146. G. NHMW 1858/0018/0146. H. NHMW 1858/0018/0146. I–J. Allmonia funiculata (Borson, 1821), Torino Hills (Italy). I. NHMW 1862/0049/0026. J. NHMW 1854/0016/0209.