Ptychidia austrorotundata, Handmann, 1882

Harzhauser, Mathias & Landau, Bernard, 2019, Turritellidae (Gastropoda) of the Miocene Paratethys Sea with considerations about turritellid genera, Zootaxa 4681 (1), pp. 1-136 : 73-74

Ptychidia austrorotundata new nom.

Figs 6Z , 19K, 19L

Turritella turris Bast. var. rotundata— Schaffer, 1912: 159, pl. 52, figs 12–14 [non Turritella rotundata Grzybowski, 1899 ].

Turritella (Haustator) eryna rotundata Schaffer — Sieber, 1960: 236, pl. 1, figs 14–15 [non Turritella rotundata Grzybowski, 1899 ].

Turritella (Haustator) eryna rotundata Schaffer, 1912 — Steininger et al., 1971: 371, pl. 6, figs 6, 9 [non Turritella rotundata Grzybowski, 1899 ].

Turritella Erynella [sic] eryna rotundata Schaffer — Stürmer, 1989: 109, pl. 3, figs 2–3, 6, 8, pl. 9, fig. 4 [non Turritella rotundata Grzybowski, 1899 ].

? Turritella eryna rotundata Schaffer — Kókay, 2002: 372, pl. 2, figs 7–11 [non Turritella rotundata Grzybowski, 1899 ].

non Turritella (Haustator) eryna cf. rotundata Schaffer, 1912 — Mikuž, 1982: 69, pl. 2, figs 2–3 [= Ptychidia vindobonensis ( Handmann, 1882)]. non Turritella (Haustator) eryna cf. rotundata Schaffer, 1912 — Mikuž, 2009: 9, pl. 1, fig. 7 [= Ptychidia vindobonensis (Hand- mann, 1882)].

Type material. The syntypes of Turritella turris var. rotundata Schaffer, 1912 are stored in the Krahuletz Museum in Eggenburg ( Austria); Eggenburgian (early Miocene); Maria Dreieichen, Eggenburg ( Austria). Herein, we designate specimen Inv. no. KM F/1152 illustrated in Schaffer (1912, pl. 52, fig. 12) as holotype of Ptychidia austrorotundata new name.

Studied material. 1 spec. KM F/1152, Maria Dreieichen ( Austria), illustrated in Schaffer (1912, pl. 52, fig. 12) ; 2 spec. NHMW 1963/0555/0000, Maria Dreieichen and Eggenburg ( Austria), illustrated in Sieber (1960, pl. 1, figs 14–15) .

Illustrated material. Fig. 19K : KM F/1152: SL: 43.5 mm, MD: 17.5 mm, illustrated in Schaffer (1912, pl. 52, fig. 12), Maria Dreieichen ( Austria); Fig. 19L : NHMW 1963/0555/0000, SL: 18.6 mm, MD: 7.9 mm, illustrated in Sieber (1960, pl. 1, fig. 14) as Turritella (Haustator) eryna rotundata Schaffer, 1912 , Maria Dreieichen ( Austria);

Revised description. Shell medium-sized, moderately slender, robust. Protoconch and early teleoconch whorls unknown. First preserved teleoconch whorl keeled at mid-whorl, coinciding with prominent, convex B spiral cord; A and C spiral cords slightly weaker, accompanied by delicate t and d cords. Primary spiral cords grading into broad, convex cords on later teleoconch whorls, two broad but less prominent secondary cords adapically to A; two to three delicate spiral cords in interspaces between A/B and B/C; d spiral cord moderately prominent, coinciding with marked angulation at transition into nearly flat base. Whorl profile changing from angulate on early spire to convex on late whorls; periphery at B spiral cord, slightly below mid-whorl. Base covered by about 9 broad spiral cords; aperture largely destroyed in available specimens. Inner lip narrowly reflected passing into narrow parietal callus. Lateral sinus with moderately steep angle, ranging around 16°; moderately deep, wide, with faint adapical and abapical inflection points. Basal sinus not visible. No information on presence of inner lirae.

Shell measurements and ratios. Only fragments are available; largest fragment ( SL: 43.5 mm, MD: 17.5 mm) suggest a total height of about 78 mm and 19 mm diameter. AA°: 20°, PA°: 14°; Lateral sinus (n = 1): LS angle = ~16°, LS p = 1.4, LS d = 5.4.

Discussion. Schaffer (1912) introduced Turritella turris var. rotundata for this early Miocene species, but was not aware that Turritella rotundata was preoccupied by T. rotundata Grzybowski, 1899 from the Neogene of Peru. Therefore, we propose austrorotundata as new name (austro = abbreviation for Austria, rotundata = as reference to Schaffer’s original name).

This species was discussed by Schaffer (1912), Sieber (1960), and Stürmer (1989) as a subspecies of Ptychidia turris (de Basterot, 1825) . Although the spiral sculpture of early teleoconch whorls is similar in both taxa, the broad, strongly convex, ‘ vermicularis -like’ spiral cords are very untypical for P. turris . In addition, Ptychidia austrorotundata is more robust and larger.

Paleoenvironment. The occurrences in the Eggenburg region in Lower Austria) all derive from coastal, sandy lagoonal and inner neritic settings.

Distribution. Western and Central Paratethys. Eggenburgian/Ottnangian (early Miocene): North Alpine -Carpathian Foreland Basin: Wirtatobel at Bregenz ( Austria), Fels/Wagram, Eggenburg, Maigen, Maria Dreieichen, Gauderndorf ( Austria) (Sieber 1969; Steininger et al. 1971); Horná Nitra Basin: Vélka Čausa ( Slovak Republic) ( Steininger et al. 1971).

Figures

FIGURE 6. Growth line traces of Turritellidae species based on own data; bold: type species of genera; black: Paratethyan species; asterisk: non-Paratethyan species (unknown basal sinuses are left blank).

FIGURE 19. A–F. Ptychidia partschi (Rolle, 1856). A1–A2. Pöls (Austria), NHMW 1861/0001/0270. B1–B2. Pöls (Austria), NHMW 2018/0275/0001, C1–C2. Holotype of Turritella (Haustator) partschi quadricarinata Sieber, 1960, Wetzelsdorf- Kreuzschaller (Austria), UMJG&P 60.658. D1–D2. Wetzelsdorf-Grössl (Austria), UMJG&P 60.632. E. Ptychidia partschi (Rolle, 1856), Wetzelsdorf-Wenzelsteffi (Austria), UMJG&P 60.847. F. Pöls (Austria), NHMW 1861/0001/0270. G–J. Ptychidia sarmatica (Papp, 1954), Hollabrunn-Aspersdorf (Austria). G. Holotype, NHMW 2018/0281/0003. H. NHMW 2018/0281/0001. I. NHMW 2013/0284/0014. J. NHMW 2018/0281/0002. K–L. Ptychidia austrorotundata new nom., Maria Dreieichen (Austria). K. KM F/1152. L. NHMW 1963/0555/0000. M–N. Ptychidia ernesti (Handmann, 1882), Steinebrunn (Austria). M1–M2. NHMW 1963/0547/0000. N. Steinebrunn (Austria), NHMW 2018/0194/0005.