Allmonia cathedralis ( Brongniart, 1823 ) Harzhauser & Landau, 2019

Harzhauser, Mathias & Landau, Bernard, 2019, Turritellidae (Gastropoda) of the Miocene Paratethys Sea with considerations about turritellid genera, Zootaxa 4681 (1), pp. 1-136 : 96-97

Allmonia cathedralis ( Brongniart, 1823) new comb.

Figs 6 Aa, 24A, 24B, 24C, 25F 1 –F 2, 25G, 25H

* Turritella cathedralis A. Br. — Brongniart, 1823: 55 (pars, French specimens only), pl. 4, fig. 6.

Turritella proto de Basterot, 1825: 30 , pl. 1 fig. 7.

Turritella quadriplicata de Basterot, 1825: 29 , pl. 1, fig. 13.

Turritella cathedralis var. C. obelisca Grateloup, 1846 : pl. 16, fig. 4.

Turritella proto Defrance — Deshayes, 1850: 42, pl. 69, figs 15–16.

Turritella cathedralis A. Brongniart, 1823 — Cossmann, 1910b: 187, figs 6, H–Ha, C 1 –C 3 [non figs C–Ca = Allmonia funiculata ( Borson, 1821) ].

[ Protoma s.s.] cathedralis Brongn. — Cossmann, 1912: 127, pl. 9, figs 3–5.

Protoma cathedralis Brongniart — Cossmann & Peyrot, 1922: 53, nr. 437, pl.2, fig. 12.

Protoma obeliscus Grateloup — Cossmann & Peyrot, 1922: 55, nr. 438, pl. 1, figs 36–37.

Protoma quadriplicata (Basterot) — Cossmann & Peyrot, 1922: 57, nr. 439 (non 429), pl. 2, fig. 7.

Protoma proto Basterot — Cossmann & Peyrot, 1922: 59, nr. 441, pl. 2, figs 5–6.

Protoma cathedralis ( Brongniart, 1823) — Lozouet et al., 2001: 28, pl. 7, figs 4a–4b, 5a–5b.

non Protoma cathedralis (Basterot) — Brébion, 1970: 5, pl. 1, fig. 2.

non Protoma cf. cathedralis (Basterot) — Brébion, 1983: 164.

Type material. Cossmann (1910b) erroneously designated a holotype from a syntype series from the Burdigalian of Léognan at Bordeaux in France ( Cossmann 1910b: 187, figs H–Ha). Thus, this specimen is the lectotype by fixation ( ICZN Art 74.6). The paralectotypes are stored in the Muséum National d’Histoire Naturelle Paris under the inventory number MNHN.F. J04336 (pers. comm. Jean-Michelle Pacaud, July 25 th, 2018).

Studied material. 5 spec. NHMW 1858/0018/0146, 5 spec. NHMW 1851/0017/1228, 5 spec. NHMW 1850/0035/0128, Léognan ( France) ; 3 spec. NHMW 1856/0035/0347, Saucats ( France) ; 1 spec. NHMW 1836/0012/0494, 3 spec. NHMW 1847/0033/0095, 7 spec. NHMW 1851/0017/1229, Bordeaux ( France) .

Illustrated material. Figs 25F 1 –F 2 : SL: 114.0 mm, MD: 29.8 mm; Fig. 25G : SL: 101.9 mm, MD: 27.7 mm; Fig. 25H : SL: 105.4 mm, MD: 28.5 mm, all from Léognan ( France), lot NHMW 1858/0018/0146.

Discussion. Herein, we designate Turritella cathedralis Brongniart, 1823 as type species of Allmonia . The status of this species, however, needs clarification. It was first described by Brongniart (1823: 55, pl. 4, fig. 6). He based his Latin descriptions on specimens from the early Miocene of the Torino Hills in northern Italy but also from the early Miocene of Léognan at Bordeaux in France, from where his illustrated specimen derived. Cossmann (1912) and Cossmann & Peyrot (1922) pointed out that cathedralis should be restricted to the French specimens. We follow this view and thus Allmonia cathedralis is a species restricted to the Miocene of the north-western Atlantic, recorded from the Aquitanian and Burdigalian of the Aquitaine Basin.

Distribution. Northeastern Atlantic. Aquitanian and Burdigalian: e.g. Léognan at Bordeaux, Saucats, Saint- Paul-lès-Dax ( France), St-Jean-de-Marsacq ( Cossmann & Peyrot 1922; Lozouet et al. 2001). Dartevelle et al. (1954) describe ‘ Protoma obeliscus ’ also from the Burdigalian of Angola, but the preservation of the illustrated specimen does not allow an identification.

Brébion (1970, 1983) described this species also from the Miocene of Angola. The illustrated specimen (MNHN. F.R06826), however, has little in common with Allmonia cathedralis and is not even a Protominae . The specimen displays a very blunt shell with an irregular angulation below the adapical suture and forms irregular humps, which are unknown from any extant Turritellidae genus.

Brébion (1964: 203, pl. 5, fig. 11) in his unpublished thesis on the ‘Redonian’ of France illustrated a single fragment identified as Protoma quadriplicata de Basterot, 1825 from the upper Miocene Tortonian of Sceaux-d’Anjou, NW France. The figured fragment does not seem to represent that species and the record could not be confirmed by Landau et al., 2018: 342).

Figures

FIGURE 1. Paleogeography of the Paratethys during the early Miocene (Burdigalian, Eggenburgian, Sakaraulian) and the middle Miocene (Langhian, Badenian, Tarkhanian) (modified after Popov et al. 2004 and Neubauer et al. 2015).

FIGURE 2. Miocene chronostratigraphy after Gradstein et al. (2012) with regional Paratethys stages after Popov et al. (2004) and Piller et al. (2007).

FIGURE 6. Growth line traces of Turritellidae species based on own data; bold: type species of genera; black: Paratethyan species; asterisk: non-Paratethyan species (unknown basal sinuses are left blank).

FIGURE 25. A–D. Allmonia paucicincta (Sacco, 1895), Burgschleinitz (Austria). A. Holotype, NHMW 1846/0010/0014. B1–B3. Collection Anton Breitenberger. C. NHMW 2018/0128/0002. D. NHMW 2018/0128/000. E. Turritella (Protoma) telleri Hilber, 1993, holotype, Ivanji dol (Slovenia), UMJG&P 1247. F–H. Allmonia cathedralis (Brongniart, 1823), Léognan (France). F1–F2. NHMW 1858/0018/0146. G. NHMW 1858/0018/0146. H. NHMW 1858/0018/0146. I–J. Allmonia funiculata (Borson, 1821), Torino Hills (Italy). I. NHMW 1862/0049/0026. J. NHMW 1854/0016/0209.