Archimediella hoernesi (Rolle, 1856) Harzhauser & Landau, 2019

Harzhauser, Mathias & Landau, Bernard, 2019, Turritellidae (Gastropoda) of the Miocene Paratethys Sea with considerations about turritellid genera, Zootaxa 4681 (1), pp. 1-136 : 29-30

Archimediella hoernesi (Rolle, 1856) new comb.

Figs 6I , 7 D–E, 10G 1 –G 2, H 1 –H 2, 10I, 10J

* Turritella Hoernesi Rolle — Rolle, 1856a: 192.

Turritella Hoernesi Rolle — Anonymous, 1856: 623.

Turritella Hoernesi Rolle n. sp. — Rolle, 1856b: 573.

Turritella Hoernesi Rolle — Hilber, 1879: 445, pl. 4, figs 12a–12c.

Turritella (Haustator) hoernesi Rolle — Sieber, 1960: 245, pl. 1, fig. 34, pl. 3, fig. 8/11.

non Turritella Hörnesi Neugeboren — Neugeboren, 1856: 97 [junior homonym].

non Turritella Hoernesi Vilanova — Vilanova y Piera, 1863, plate captions, pl. 8, fig. 24 [junior homonym].

non Turritella Hoernesi Micht. in coll. — Manzoni, 1869: 496, pl. 2, fig. 12 [junior homonym].

non Turritella (Haustator) hoernesi Rolle, 1856 — Katona, et al. 2011: 7, pl. 1, fig. 6 [probably an Allmonia species].

Type material. The syntypes of Rolle (1856a) could not be located. As they are not stored in the Geological Survey in Vienna, the specimens are most probably lost. Therefore we designate specimen UMJG&P 74.936 ( Figs 10H 1–H 2 ) from Wetzelsdorf-Winkeltoni ( Austria) as neotype .

Studied material. 11 spec. NHMW 1858/0020/0024, Guglitz at St. Florian ( Austria) ; 1 spec. NHMW 1989/0030/0001, Wetzelsdorf ( Austria) ; 1 spec. UMJG&P 61.170, Wetzelsdorf-Kreuzschaller ( Austria) , illustrat- ed in Sieber (1960, pl. 1, fig. 34); Wetzelsdorf-Kreuzschaller ( Austria) , 35 spec. UMJG&P 60.895, Wetzelsdorf- Kreuzschaller ( Austria) ; 65 spec. UMJG&P 60.894, 1 spec. UMJG&P 74.936-1, 1 spec. UMJG&P 74.936-2, 58 spec. UMJG&P 74.936, Wetzelsdorf-Winkeltoni ( Austria) ; 2 spec. NHMW 2018/0308/0001– NHMW 2018/0308/0002, Wetzelsdorf ( Austria) .

Illustrated material. Fig 7D : NHMW 2018/0308/0001, Fig 7E : NHMW 2018/0308/0001, both Wetzelsdorf ( Austria); Figs 10G 1 –G 2 : UMJG&P 61.170, SL: 40.4 mm, MD: 10.6 mm, Wetzelsdorf-Kreuzschaller ( Austria), illustrated in Sieber (1960, pl. 1, fig. 34); Figs 10H 1 –H 2 : UMJG&P 74.936-1 (neotype), SL = 35.8 mm, MD = 8.0 mm, Wetzelsdorf-Winkeltoni ( Austria) ; Fig. 10I : UMJG&P 74.936-2, SL = 41.0 mm, MD = 10.4 mm, Wetzelsdorf- Winkeltoni ( Austria) ; Fig. 10J : NHMW 1858/0020/0024, SL: 13.5 mm, MD: 4.3 mm, Guglitz ( Austria).

Revised description. Shell small, slender to moderately slender, of ca. 16 teleoconch whorls. Protoconch only partly preserved; second protoconch whorl moderately high with mid-whorl angulation. Neanic whorls campanulate with low-angled, concave sutural ramp and prominent keel formed by B spiral cord, accompanied by weaker C spiral cord. Faint peribasal d spiral cord may appear on neanic whorls. Numerous, densely spaced, delicate secondary spiral cords with convex tops present throughout ontogeny. A spiral cord successively increasing in strength between 9 th to 11 th teleoconch whorls, resulting in tricostate sculpture of primary spiral cords with convex tops. B and C primaries always more prominent interspace between both cords slightly widening during growth. Whorl profile weakly convex to nearly flat sided with moderately incised suture. Faint d spiral cord visible on last whorl, coinciding with transition into nearly flat base. Lateral sinus angle moderately steep; moderately deep with faint adapical inflection points; adapical LS p. Basal sinus nearly straight to weakly opisthocyrt. Base (of subadult specimens) moderately convex, covered by about five secondary spiral cords and delicate tertiary spiral cords in interspaces. Aperture missing in all specimens. Inner lirae comprising about seven prominent palatal lirae and one strong basal spiral cord.

Shell measurements and ratios. All specimens are incomplete and the last whorl is missing in all large specimens; largest fragments range around 41 mm suggesting a total height of c. 46 mm and c. 12 mm diameter. AA (n=10): 19.0° (σ = 1.7°), PA: 13.6° (σ = 0.8°). Lateral sinus (n = 5): LS angle = 14.8° (σ = 3.6°), LS p = 0.9 (σ = 0.1), LS d = 5.6 (σ = 0.4).

Discussion. Turritella hoernesi has been repeatedly used as a name for a Neogene species. Rolle (1856) introduced the name in the same year as Neugeboren (1856), who proposed Turritella Hörnesi as new name for a taxon from the Badenian (middle Miocene) of Lăpugiu de Sus in Romania, but it was not made clear to which species he was referring. The species has never been illustrated and Rolle’s specimens could not be located. From the description it is clear that Neugeboren (1856) described shells related or conspecific with Ptychidia vindobonensis Handmann, 1882 (= Turritella turris sensu Hörnes, 1855 ). Thus, Turritella hoernesi Rolle and T. hoernesi Neugeboren are homonyms, but certainly not synonyms. The paper by Rolle was presented on February 26 th, 1856 in the ‘Sitzungen der k.k Reichsanstalt’ (= conference or meeting), but the publication date is not provided in the volume. The reprint of the description by Rolle (1856a), however, is given in the issue from March 1 st, 1856 in the ‘ Oesterreichisch Kaiserliche Wiener Zeitung ’, an important Austrian newspaper at that time ( Anonymous 1856). Therefore, the description of Turritella hoernesi Rolle, 1856 was published simultaneously with the presentation in April 1856. The paper by Neugeboren is indicated to have been published in July 1856 (see Verhandlungen und Mitteilungen des Siebenbürgischen Vereins für Naturwissenschaften zu Hermannstadt, 1856, 7: 101). Therefore, T. hoernesi Rolle gains priority. More than a decade later, Manzoni (1869) described another species as Turritella hoernesi from the Italian Neogene. The illustration of this species in Manzoni (1869) suggests that it might represent a fragment of an Allmonia species.

Paleoenvironment. The species is recorded so far only from marly clay of Wetzelsdorf and Pöls in the Styrian Basin. Especially Wetzelsdorf is known for the frequent occurrence of the rostellariid Tibia, which suggests deeper water conditions ( Kronenberg 2008).

Distribution. Central Paratethys. Badenian (middle Miocene): Styrian Basin: Wetzelsdorf at Graz, Pöls ( Austria).

Figures

FIGURE 1. Paleogeography of the Paratethys during the early Miocene (Burdigalian, Eggenburgian, Sakaraulian) and the middle Miocene (Langhian, Badenian, Tarkhanian) (modified after Popov et al. 2004 and Neubauer et al. 2015).

FIGURE 2. Miocene chronostratigraphy after Gradstein et al. (2012) with regional Paratethys stages after Popov et al. (2004) and Piller et al. (2007).

FIGURE 6. Growth line traces of Turritellidae species based on own data; bold: type species of genera; black: Paratethyan species; asterisk: non-Paratethyan species (unknown basal sinuses are left blank).

FIGURE 7. Internal lirae in selected Turritellidae species. A. Archimediella abundans (Handmann, 1882), NHMW 2018/0307/0001, Coşteiu de Sus (Romania). B. Archimediella abundans (Handmann, 1882), NHMW 2018/0307/0002, Coşteiu de Sus (Romania). C. Archimediella carpathica new sp., NHMW 2018/0272/0006, Gainfarn (Austria). D. Archimediella hoernesi (Rolle, 1856), NHMW 2018/0308/0001, Wetzelsdorf (Austria). E. Archimediella hoernesi (Rolle, 1856), NHMW 2018/0308/0002, Wetzelsdorf (Austria). F. Helminthia vermicularis (Brocchi, 1814), NHMW 2018/0272/0007, Gainfarn (Austria). G. Helminthia vermicularis (Brocchi, 1814), NHMW 2018/0181/0010, Lăpugiu de Sus (Romania). H. Oligodia bicarinata (Eichwald, 1830), NHMW 2018/0272/0008, Gainfarn (Austria); I. Oligodia spirata (Brocchi, 1814), NHMW 2018/0272/0009, Gainfarn (Austria). J. Ptychidia vindobonensis (Handmann, 1882), NHMW 2018/0181/0011, Lăpugiu de Sus (Romania).

FIGURE 10. A–B. Archimediella cochlias (Bayan, 1873). A1–A4. Sangonini di Lugo (Italy) GBA 2010/152/0055. B. Gnata di Salcedo, GBA 2010/148/0062. C–F. Archimediella abundans (Handmann, 1882). C. Steinebrunn (Austria), NHMW 1846/0037/0393. D1–D2. Steinebrunn (Austria), NHMW 1871/0010/0382. E1–E2. Lăpugiu de Sus (Romania), NHMW 1869/0019/0204. F1–F2. Steinebrunn (Austria), NHMW 1963/0518/0000. G–J. Archimediella hoernesi (Rolle, 1856). G1–G2. Wetzelsdorf-Kreuzschaller (Austria), UMJG&P 61.170. H1–H2. Neotype, Wetzelsdorf-Winkeltoni (Austria), UMJG&P 74.936- 1. I. Wetzelsdorf-Winkeltoni (Austria), UMJG&P 74.936-2. J. Guglitz (Austria), NHMW 1858/0020/0024.