Archimediella carpathica, Harzhauser & Landau, 2019

Harzhauser, Mathias & Landau, Bernard, 2019, Turritellidae (Gastropoda) of the Miocene Paratethys Sea with considerations about turritellid genera, Zootaxa 4681 (1), pp. 1-136 : 30-34

Archimediella carpathica new sp.

Figs 6H , 7C , 11A 1 –A 2 , 11B, 11C, 11D, 11E, 11F, 11G 1 –G 2

Turritella Archimedis Brong. — Hörnes, 1855: 424 (pars) [non Turritella archimedis Brongniart, 1823 ].

[ Turritella (Oligodia) Archimedis Brong. ] var. tricarinata— Handmann, 1882: 217 [non Turritella tricarinata ( Brocchi, 1814) , nec Turritella tricarinata King, 1832 ].

? Turritella Sokolovi n. sp. — Mikhailovsky, 1903: 52, 215, pl. 3, figs 4–6.

Turritella dertonensis Mayer — Friedberg, 1909: 256, pl. 2, fig. 13 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella dertonensis var. subconica Sacco — Friedberg, 1909: 257, pl. 2, fig. 14 [non Archimediella dertonensis subconica Sacco, 1895 ].

Turritella dertonensis May. — Friedberg, 1914: 338 , pl. 19, fig. 25 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella dertonensis May. var. subconica Sacco (?) — Friedberg, 1914: 339 , pl. 19, fig. 26 [non Archimediella dertonensis subconica Sacco, 1895 ].

Turritella communis Risso var. tricarinata Q. — Quitzow, 1922: 17 , pl. 1, fig. 16 [non Turritella tricarinata ( Brocchi, 1814) ].

Turritella (Haustator) pythagoraica (HiIb.) — Sieber, 1953: 185 [non Archimediella pythagoraica ( Hilber, 1882) ].

? Turritella (Archimediella) pythagoraica Hilber, 1882 — Moisescu, 1955a: 124, pl. 10, figs 14–15 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella (Archimediella) dertonensis May. — Csepreghy-Meznerics, 1956: 383 , pl. 2, fig. 18 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella (Archimediella) dertonensis subconica Sacco — Csepreghy-Meznerics, 1956: 383, pl. 2, fig. 28 [non Archimediella dertonensis subconica Sacco, 1895 ].

Turritella (Archimediella) dertonensis May. — Sieber, 1947: 156 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella (Haustator) pythagoraica Hilb. — Sieber, 1956: 238 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella (Archimediella) pythagoraica pythagoraica Hilber — Sieber, 1960: 254, pl. 2, figs 6, 9, 14, pl. 3, fig. 8/24 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella (Archimediella) erronea subpythagoraica Friedbg. — Sieber, 1960: 254, pl. 2, fig. 15 [non Turritella erronea var. subpythagoraica Friedberg, 1914 ].

Turritella (Archimediella) dertonensis dertonensis Mayer — Sieber, 1960: 255, pl. 2, figs 11–12, pl. 3, fig. 8/25 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella (Archimediella) dertonensis subconica Sacco — Sieber, 1960: 256, pl. 2, fig. 18 [non Archimediella dertonensis subconica Sacco, 1895 ].

Turritella (Archimediella) dertonensis persulcata Sacco — Sieber, 1960: 256, pl. 2, fig. 10.

Turritella (Archimediella) dertonensis Mayer — Marinescu, 1961: 529, pl. 4, fig. 18 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella (Archimediella) pythagoraica pythagoraica Hilber — Švagrovský, 1960: 87, pl. 9, figs 1–3 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella pythagoraica Hilb. — Švagrovský, 1964: 84, pl. 17, fig. 5 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella cfr. pythagoraica Hilber, 1882 — Strausz, 1966: 90, pl. 2, fig. 18 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella benoisti Cossmann & Peyrot, 1921 — Strausz, 1966: 91, pl. 2, figs 19–20 [non Archimediella benoisti ( Cossmann & Peyrot, 1922) ].

Turritella (Archimediella) dertonensis Mayer — Kókay, 1966: 39, pl. 3, fig. 13 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella erronea dertornatior [sic] Sacco — Krach, 1967: 224 [non Archimediella dertornatior Sacco, 1895 ].

Turritella (Archimediella) dertonensis Mayer, 1868 — Hinculov, 1968: 133, pl. 31, figs 18–19 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella (Archimediella) pythagoraica Hilber, 1882 — Hinculov, 1968: 133, pl. 32, figs 1–2 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella benoisti (Cossmann et Peyrot) — Stancu et al., 1971: 125, pl. 8, fig. 2 [non Turritella erronea var. subpythagoraica Friedberg, 1914 ].

? Turritella benoisti Cossmann — Peyrot, 1924 — Nicorici & Sagatovici, 1973: 163, pl. 23, figs 11–14 [non Turritella erronea var. subpythagoraica Friedberg, 1914 ].

Turritella pythagoraica Hilber, 1882 — Nicorici & Sagatovici, 1973: 164, pl. 23, figs 15–16 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella (Archimediella) dertonensis Mayer, 1868 — Bałuk, 1975: 112, pl. 12, figs 10–12 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella (Archimediella) pythagoraica pythagoraica Hilber, 1882 — Steininger et al., 1978: 332, pl. 2, figs 6–8 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella (Archimediella) dertonensis Mayer, 1868 — Steininger et al., 1978: 332, pl. 2, figs 9–10 [non Archimediella dertonensis ( Mayer, 1868) ].

Turritella (Archimediella) pythagoraica Hilber (1882) — Švagrovský, 1981: 128, pl. 40, figs 1–2 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella (Archimediella) erronea subpythagoraica Friedberg, 1914 — Švagrovský, 1982: 16, pl. 4, fig. 5 [non Turritella erronea var. subpythagoraica Friedberg, 1914 ].

Turritella (Archimediella) dertonensis dertonensis Mayer, 1868 — Švagrovský, 1982: 15, pl. 4, fig. 7 [non Archimediella dertonensis ( Mayer, 1868) ].

[ Archimediella archimedis ( Brongniart, 1823) ] var. dertornatior Sacco, 1895 — Ferrero Mortara et al., 1984: 229.

Turritella (Arcimediella [sic]) dertonensis Mayer, 1868 — Atanacković, 1985: 93, pl. 21, figs 18–20 [non Archimediella dertonensis ( Mayer, 1868) ].

Archimediella Archimediella [sic] pythagoraica pythagoraica Hilber — Stürmer, 1989: 146, pl. 6, figs 7–9, 11–12, pl. 11, figs 6–7, pl. 12, figs 7, 14 [non Archimediella pythagoraica ( Hilber, 1882) ].

Archimediella Archimediella [sic] erronea persulcata Sacco — Stürmer, 1989: 135, pl. 6, figs 6, 10, pl. 11, fig. 11.

Turritella pythagoraica Hilber — Ionesi & Nicorici, 1994: 60, pl. 4, figs 3–6 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella (Archimediella) archimedis mioconica Sacco — Kókay, 1996: 455, pl. 1, fig. 3 [non Archimediella mioconica Sacco, 1895 ].

Turritella (Archimediella) pythagoraica pythagoraica Hilber — Schultz, 1998: 56, pl. 21, fig. 12 [non Archimediella pythagoraica ( Hilber, 1882) ].

Turritella (Archimediella) dertonensis dertonensis Mayer — Schultz, 1998: 56, pl. 21, fig. 13 [non Archimediella dertonensis ( Mayer, 1868) ].

? Turritella (Archimediella) erronea subpythagoraica Friedberg, 1914 — Katona et al., 2011: 7, pl. 2, fig. 1 [non Turritella erronea var. subpythagoraica Friedberg, 1914 ].

Turritella pythagoraica pythagoraica Hilber — Hladilová & Fordinál, 2013: 39, fig. 5a [non Archimediella pythagoraica ( Hilber, 1882) ].

Oligodia pythagoraica ( Hilber, 1882) — Harzhauser et al., 2018: 160, figs 10G–H [non Archimediella pythagoraica ( Hilber, 1882) ].

Types. Holotype NHMW 2018/0272/002 ( Figs 11A 1 –A 2 ), SL: 60.4 mm, MD: 14.9 mm, Gainfarn ( Austria) . Paratypes NHMW 2018/0272/0003 ( Fig. 11B ), SL: 58.0 mm, MD: 13.6 mm, Gainfarn ( Austria) ; NHMW 2018/0271/0004 ( Fig. 11C ), SL: 53.5 mm, MD: 13.5 mm, Enzesfeld ( Austria); NHMW 2018/0271/0005 ( Fig. 11D ), SL: 56.0 mm, MD: 12.6 mm, Enzesfeld ( Austria); NHMW 2018/0272/0004 ( Fig. 11E ), SL: 50.3 mm, MD: 14.5 mm, Gainfarn ( Austria); NHMW 1963/0550/0000 ( Fig. 11F ), Braunsdorf ( Austria), illustrated in Sieber (1960, pl. 2, fig. 14) as Turritella (Archimediella) pythagoraica Hilber ; NHMW 2018/0274/0001 Figs 11G 1 –G 2 ), SL: 4.3 mm, MD: 1.7 mm, Bernhardsthal ( Austria).

Type stratum. Marly sand of the Jakubov Formation of the Baden Group ( Zuschin et al. 2007).

Type locality. Gainfarn ( Austria) .

Age. Middle Miocene, Badenian (= late Langhian).

Etymology. Referring to the Carpathian Mountains, because this species is found in the basins surrounding this mountain chain.

Additional material. 1 spec. NHMW 1963/0545/0000, Enzesfeld ( Austria) , illustrated in Sieber (1960, pl. 2, fig. 6) as Turritella (Archimediella) pythagoraica Hilber ; 1 spec. NHMW 1963/0548/0000, Enzesfeld ( Austria) , illustrated in Sieber (1960, pl. 2, fig. 15) as Turritella (Archimediella) erronea subpythagoraica Friedberg ; 3 spec. NHMW 2018/0271/0001, 1 spec. NHMW 2018/0271/0004, 1 spec. NHMW 2018/0271/0005, 3 spec. NHMW 2018/0271/0002, 30 spec. NHMW 2018/0271/0003, Enzesfeld ( Austria) ; 27 spec. NHMW 1989/0037/0008, 1 spec. NHMW 2018/0272/0002, 1 spec. NHMW 2018/0272/0003, 1 spec. NHMW 2018/0272/0004, 11 spec. NHMW 2018/0273/0001, 16 spec. NHMW 2018/0273/0002, Vienna-Grinzing ( Austria) ; 1 spec. NHMW 1963/0511/0000, Vienna-Ottakring ( Austria) , illustrated in Sieber (1960, pl. 2, fig. 9) as Turritella (Archimediella) pythagoraica Hilber ; 1 spec. NHMW 1963/0550/0000, Braunsdorf ( Austria) , illustrated in Sieber (1960, pl. 2, fig. 14) as Turritella (Archimediella) pythagoraica Hilber ; 1 spec. NHMW 1963/0553/0000, Möllersdorf ( Austria) , illustrated in Sieber (1960, pl. 2, fig. 11) as Turritella (Archimediella) dertonensis Mayer ; 1 spec. NHMW 1963/0544/0000, Windpassing ( Austria) , illustrated in Sieber (1960, pl. 2, fig. 12) Turritella (Archimediella) dertonensis Mayer ; 1 spec. NHMW 1963/0516/0000, Ottakring ( Austria) , illustrated in Sieber (1960, pl. 2, fig. 10) Turritella (Archimediella) dertonensis persulcata Sacco ; 1 spec. NHMW 1963/0533/0000, Ottakring ( Austria) , illustrated in Sieber (1960, pl. 2, fig. 18) Turritella (Archimediella) dertonensis subconica Sacco ; 1 spec. NHMW 1859/0038/0021, 1 spec. NHMW 1859/0038/0001, Vienna-Speising ( Austria) ; 1 spec. NHMW 2018/0274/0001, Bernhardsthal , OMV drilling, Be 6, depth: 1397–1406 m ( Austria) ; 1 spec. NHMW 2018/0272/0006, Gainfarn ( Austria) ( Fig. 7C ).

Description. Shell medium-sized, slender, comprising up to 20 teleoconch whorls. Protoconch moderately high, consisting of less than two strongly convex whorls with bulbous initial part; diameter 290 μm. Transition into teleoconch marked by onset of keel-like B spiral cord and slightly weaker C spiral cord. Both primary spiral cords persist throughout ontogeny forming two moderately raised cords with convex tops, separated by flat to weakly concave interspace. Sutural ramp indistinct, flat to feebly convex on late whorls. Weakly raised A spiral cord may develop on sutural ramp, typically accompanied adapically by two secondary spiral cords and one abapical one. Weaker tertiary spiral cords may appear in interspaces. Three tertiary spiral cords below C primary cord. Suture moderately to distinctly incised; whorl profile shifting from weakly convex to moderately frustate during ontogeny. Lateral sinus with moderately steep angle, ranging around 18.5°; moderately deep with weak adapical inflection point; occasionally, weak abapical inflection point appears close to suture; abapical LS p. Basal sinus weakly opisthocyrt to slightly sigmoidal. Aperture semicircular; columella concave; parietal callus glossy, thin, weakly expanding. Base moderately convex with about five to eight wide-spaced spiral cords and intercalated secondary cords of variable strength. Inner lirae present on late teleoconch whorls, consisting of eight distinct palatal lirae and one prominent basal cord in abaxial half accompanied by 3–4 weaker spiral cords.

Shell measurements and ratios. All specimens are incomplete; the SL/MD ratio of rather complete specimens ranges around 4.0. Based on this ratio, the largest fragments containing the last whorl suggest a maximum SL of 68 to78 mm and a maximum diameter of 16.9 to 19.6 mm. AA (n=12): 19.7° (σ = 3.2°), PA: 12.6° (σ = 0.9°). Lateral sinus (n = 8): LS angle = 18.5° (σ = 3.0°), LS p = 1.1 (σ = 0.1), LS d = 5.4 (σ = 1.1).

Discussion. The species displays moderate variability concerning the strength of the two primary spiral cords but develops a broad range of morphotypes with variable strength and number of secondary and tertiary spiral cords. In some specimens, these spirals attain about the same strength, forming a regular sculpture and even the primary cords may bear distinct secondary cords. Especially the presence and strength of the A spiral cord is highly variable ranging from nearly absent to prominent, sometimes resulting in a ‘tricarinate’ profile of late teleoconch whorls. This variability is reflected in a large number of names that have been used in the Paratethyan literature. In our opinion, however, none of these names can be used for this species. Most authors have misidentified this species as Archimediella pythagoraica ( Hilber, 1882) [= A. indigena ( Eichwald, 1830) ]. Archimediella indigena differs in its smaller size, the less defined sutural ramp, and the less prominent primary cords (of which the C spiral cord may form an abapical angulation). Judging from the short diagnosis, Handmann (1882) seems to have been the first who recognized this species as distinct taxon and proposed Turritella (Oligodia) Archimedis Brong. var. tricarinata as its name. Turritella tricarinata , however, was already preoccupied by taxa described by Brocchi (1814) and King (1832). Later, Friedberg (1909) recognized the species again and identified it as Turritella dertonensis Mayer, 1868 and T. dertonensis subconica Sacco, 1895 . These identifications were uncritically adopted by many subsequent authors (e.g. Bałuk 1975; Švagrovský 1982; Schultz 1998). Archimediella dertonensis was described by Mayer (1868: 106, pl. 2, fig. 6) from the Tortonian of Stazzano and Sassuolo in northern Italy. Archimediella dertonensis and the variety subconica Sacco, 1895 differ from A. carpathica in their smaller size, the strongly reduced second- ary sculpture and the prominent B spiral cord (see syntype of A. dertonensis , Fig. 12 C : BS.061.03.004, SL: 27.0 mm, MD: 9.0 mm, Stazzano ( Italy), illustrated in Sacco (1895, pl. 1, fig. 39). Archimediella persulcata Sacco, 1895 (introduced as variety of dertonensis ) has little in common with the Paratethyan species and differs also from A. dertonensis in the close-set B and C spiral cords in the abapical half of the whorl and the very prominent, regular secondary spiral cords (see Caprotti 1975, 2011). Archimediella miotaurina Sacco 1895 , from the lower Miocene of the Torino hills, differs from A. carpathica in the keel-like B spiral cord and the much weaker C spiral cord (see Ferrero-Mortara et al. 1984); its generic placement in Archimediella might need revision. Archimediella dertornatior Sacco, 1895 , from the late Miocene of Stazzano ( Italy), is less slender and develops a distinct sutural ramp [see syntypes Fig. 12A : BS.061.03.003, SL: 41.5 mm, MD: 11.0 mm, illustrated in Sacco (1895, pl. 1, fig. 38a); Fig. 12B : BS.061.03.003bis, SL: 38.0 mm, MD: 12.0 mm, syntype illustrated in Sacco (1895, pl. 1, fig. 38b)].

Archimediella carpathica is reminiscent of A. abundans ( Handmann, 1882) in general shape and sculpture but can be separated based on its more slender profile and the less prominent B and C primary spiral cords, which have convex tops instead of forming sharp keels; moreover, the secondary spiral cords are more prominent in A. carpathica . In addition, the C spiral cord of A. carpathica is weaker on neanic whorls.

Strausz (1966) identified very slender morphotypes with prominent A spiral cord as Turritella benoisti Cossmann & Peyrot, 1922 , which was originally described from the Serravallian of Salle in France. Despite the overall similarity, the French species has higher and more convex whorls and the suture is deeper (see syntype of Turritella benoisti MNHN.F. J05849 : http://mediaphoto.mnhn.fr/media/1446626236657KcVpm2LghIcnbBzM).

Harzhauser et al. (2018) placed this species in Oligodia , but the B+C succession of primary spirals of A. carpathica differs clearly from the B–D succession of Oligodia .

Turritella sokolovi Mikhailovsky, 1903 , from the Tarkhanian (early Middle Miocene) of Tomakivka (S. Ukraine) might represent an eastern Paratethyan occurrence of A. carpathica . This name would have priority, but the fragmentary preservation does not allow a clear identification (= species inquirenda). Moreover, no types are preserved, because Mikhailovsky (1903) used only plaster casts. We were not able to locate these casts.

Paleoenvironment. The species seems to have preferred shallow marine, inner neritic and lagoonal environments, partly with sea grass meadows ( Harzhauser et al. 2018).

Distribution. Central Paratethys. Badenian, Middle Miocene: North Alpine Foreland Basin: Grund, Windpassing, Braunsdorf ( Austria) ( Sieber 1960); Carpathian Foreland Basin: Gliwice (Alt-Gleiwitz), Korytnica, Małoszów, Gartatowice, Hołubica ( Poland) ( Friedberg 1914; Quitzow 1922; Bałuk 1975); Vienna Basin: Möllersdorf, Vien- na-Ottakring, Enzesfeld, Pötzleinsdorf ( Austria) ( Sieber 1953, 1960); Borský Mikuláš, Kumice, Devínska Nová Ves ( Slovakia) ( Švagrovský 1960, 1964, 1981); Eisenstadt-Sopron Basin: Mattersburg ( Austria) ( Sieber, 1956); Oberpullendorf Basin: Ritzing ( Austria) ( Sieber 1960); Klagenfurt Basin: Mettersdorf ( Austria) ( Sieber 1960); Pannonian Basin: Illés-Street (Budapest), Hidas, Szabolcs, Pécs, Szob ( Hungary) ( Csepreghy-Meznerics 1956; Kókay 1966; Strausz 1966); southern Pannonian Basin: Hrvaćani-Drenik ( Bosnia and Herzegovina) ( Atanacković 1985); Caransebeş-Mehadia Basin: Valea Satului, Valea Bela Reca ( Romania) ( Hinculov 1968); Buzău Basin: Vale de Muscel ( Romania) ( Ionesi & Nicorici 1994); Zârand Basin: Minişul de Sus, ( Romania) ( Nicorici & Sagatovici 1973); Transylvanian Basin: Bujtur, Lăpugiu de Sus ( Moisescu 1955a; Strausz 1966). Eastern Paratethys. Turritella sokolovi Mikhailovsky, 1903 , from the Tarkhanian (early Middle Miocene) of Tomakivka (S. Ukraine) might represent an eastern Paratethyan occurrence.

Figures

FIGURE 1. Paleogeography of the Paratethys during the early Miocene (Burdigalian, Eggenburgian, Sakaraulian) and the middle Miocene (Langhian, Badenian, Tarkhanian) (modified after Popov et al. 2004 and Neubauer et al. 2015).

FIGURE 2. Miocene chronostratigraphy after Gradstein et al. (2012) with regional Paratethys stages after Popov et al. (2004) and Piller et al. (2007).

FIGURE 6. Growth line traces of Turritellidae species based on own data; bold: type species of genera; black: Paratethyan species; asterisk: non-Paratethyan species (unknown basal sinuses are left blank).

FIGURE 7. Internal lirae in selected Turritellidae species. A. Archimediella abundans (Handmann, 1882), NHMW 2018/0307/0001, Coşteiu de Sus (Romania). B. Archimediella abundans (Handmann, 1882), NHMW 2018/0307/0002, Coşteiu de Sus (Romania). C. Archimediella carpathica new sp., NHMW 2018/0272/0006, Gainfarn (Austria). D. Archimediella hoernesi (Rolle, 1856), NHMW 2018/0308/0001, Wetzelsdorf (Austria). E. Archimediella hoernesi (Rolle, 1856), NHMW 2018/0308/0002, Wetzelsdorf (Austria). F. Helminthia vermicularis (Brocchi, 1814), NHMW 2018/0272/0007, Gainfarn (Austria). G. Helminthia vermicularis (Brocchi, 1814), NHMW 2018/0181/0010, Lăpugiu de Sus (Romania). H. Oligodia bicarinata (Eichwald, 1830), NHMW 2018/0272/0008, Gainfarn (Austria); I. Oligodia spirata (Brocchi, 1814), NHMW 2018/0272/0009, Gainfarn (Austria). J. Ptychidia vindobonensis (Handmann, 1882), NHMW 2018/0181/0011, Lăpugiu de Sus (Romania).

FIGURE 11. A–G. Archimediella carpathica new sp. A1–A2. Holotype, Gainfarn (Austria), NHMW 2018/0272/002. B. Paratype, Gainfarn (Austria), NHMW 2018/0272/0003. C. Paratype, Enzesfeld (Austria), NHMW 2018/0271/0004. D. Paratype, Enzesfeld (Austria), NHMW 2018/0271/0005. E. Gainfarn (Austria), NHMW 2018/0272/0004. F. Paratype, Braunsdorf (Aus- tria), NHMW 1963/0550/0000. G1–G2. Paratype, Bernhardsthal (Austria), NHMW 2018/0274/0001. H–O. Archimediella indigena (Eichwald, 1830). H. Czepiele (Ukraine), GBA 2008/008/0001. I. Czepiele (Ukraine), GBA 2008/008/0002. J. Czepiele (Ukraine), GBA 2008/008/0003. K. Czepiele (Ukraine), GBA 2008/008/0004. L. Czepiele (Ukraine), GBA 2008/008/0005. M. Podhorce (Ukraine), NHMW 1907/0004/0017. N. Podhorce (Ukraine), NHMW 1907/0004/0017. O. Podhorce (Ukraine), NHMW 1907/0004/0017.

FIGURE 12. A–B. Archimediella dertornatior Sacco, 1895. A. Syntype, Stazzano (Italy), BS.061.03.003. 12B1–B2. Syntype, Stazzano (Italy), BS.061.03.003bis. C. Archimediella dertonensis (Mayer, 1868), Stazzano (Italy), BS.061.03.004.