Scythropopsis cornuta ( Bates, 1880 ) Santos-Silva & Botero & Wappes, 2020

Santos-Silva, Antonio, Botero, Juan Pablo & Wappes, James E., 2020, Neotropical Acanthoderini (Coleoptera, Cerambycidae, Lamiinae): Synonymies and new status in some genera, new species, transferences and new distributional records, Papéis Avulsos de Zoologia 60, pp. 1-40 : 11-14

Scythropopsis cornuta ( Bates, 1880) , comb. nov. ( Figs. 56-61 )

Acanthoderes cornutus Bates, 1880: 142 .

Acanthoderes cornuta ; Lameere, 1883: 62 (cat.); Blackwelder, 1946: 610 (checklist); Chemsak et al., 1992: 130 (checklist).

Acanthoderes (Psapharochrus) cornuta ; Aurivillius, 1923: 386 (cat.); Gilmour, 1965: 614 (cat.); Monné & Giesbert, 1994: 230 (checklist); Monné, 1994: 60 (cat.).

Psapharochrus cornuta ; Turnbow et al., 2003: 29 (distr.).

Psapharochrus cornutus ; Monné, 2005: 203 (cat.); Hovore, 2006: 376 (distr.); Monné, 2019: 293 (cat.).

Description: Male ( Figs. 57-61 ): Integument mostly dark brown, almost black on some areas; anteclypeus, apex of labrum, parts of mouthparts, and apex of palpomeres reddish-brown; antennomeres V-XI with orange ring on basal third; central area of anterior third of elytra with large, triangular reddish-brown macula.

Head: Frons, area between antennal tubercles and beginning of upper eye lobes finely, sparsely punctate; with moderately dense brownish pubescence (more reddish-brown close to eyes and postclypeus), with white pubescence interspersed. Remaining surface of vertex smooth, with dark brown pubescence centrally, partially obscuring integument, and longitudinal yellowish-white pubescent band laterally. Area behind upper eye lobes with sparse fringe of yellowish-white pubescence close to eye, glabrous on narrow sulcate area close to eye, with yellowish-brown pubescence close to sulcate area, nearly glabrous toward prothorax. Area behind lower eye lobes glabrous on sulcus adjacent to eye, with narrow (widened near inferior curvature of eye) yellowish-brown pubescent band close to sulcus (pubescence sparser than behind lower eye lobe), remaining surface glabrous. Genae almost twice length of lower eye lobe; with fringe of yellowish-brown pubescence close to eye, and sparse yellowish-brown pubescence, with white pubescence interspersed on remaining surface, narrow distal area glabrous. Postclypeus coarsely, sparsely punctate on wide central area, smooth laterally; with short, bristly reddish-brown pubescence not obscuring integument on wide central area more so on center of this area, laterally glabrous; with long, sparse, erect dark setae on wide central area. Posterior ⅔ of labrum coplanar with anteclypeus, inclined at anterior third; with yellowish-brown pubescence not obscuring integument, on posterior ⅔, anterior third nearly glabrous, anterior margin with fringe of brown setae (apex of nearly all setae yellowish); with transverse row of long, erect, dark setae near curvature between coplanar and inclined area. Gulamentum glabrous, wide posterior area smooth, except transverse striae on center of anterior region of this area; anterior area depressed, transversely striate, with sparse reddish-brown pubescence close to anterior margin. Distance between upper eye lobes 0.55 times length of scape; in frontal view, distance between lower eye lobes equal to length of scape. Antennae 1.65 times elytral length, reaching elytral apex near apex of antennomere VIII. Scape clavate, distinctly sulcate dorsally at anterior third; dorsally with reddish-brown pubescence except wide transverse white pubescent band centrally, and narrow white pubescent band at apex; remaining surface with white pubescence not obscuring integument. Pedicel with transverse white pubescent band dorsally near base, brown on remaining dorsal surface except yellowish-white on outer side close to apex; ventral and lateral surfaces with white pubescence. Antennomeres III and IV with two white pubescent rings, one basally, another after middle, ventrally fused by longitudinal band; remaining surface with brown pubescence. Antennomeres V-XI with white pubescence on basal half, brown on distal half; distal third of antennomere IX and basal ⅔ of antennomere X with long, erect dark setae. Antennal formula (ratio) based on length of antennomere III: scape = 0.73; pedicel = 0.29; IV = 0.76; V = 0.45; VI = 0.39;VII = 0.35;VIII = 0.31; IX = 0.27; X = 0.21; XI = 0.24.

Thorax: Lateral tubercle of prothorax large, conical. Pronotum with large, nearly conical tubercle each side, with wide, rounded apex; with narrow, carina-shaped tubercle centrally, from posterior sulcus to anterior margin, slightly widened posteriorly; posterior sulcus distinct laterally, nearly indistinct toward center; coarsely, sparsely punctate between tubercles, slightly coarser and more abundant between anterior margin and base of lateral tubercles,and along posterior sulcus,shallower, sparser on lateral tubercles of prothorax; with white, yellowish-brown, and pale-yellow pubescence intermixed, partially obscuring integument in some areas. Sides of prothorax coarsely, sparsely punctate (punctures distinctly coarser close to posterior margin); pubescence as on pronotum. Prosternum with pale-yellow pubescence not obscuring integument. Prosternal process with widest central area about as wide as procoxal cavity; lateral margins sinuous and elevated; central area longitudinally slightly elevated; with sparse pale-yellow pubescence. Ventral surface of meso- and metathorax with pale-yellow pubescence not obscuring integument, sparser on center of meso- and metaventrite. Mesoventral process with apex slightly wider than mesocoxal cavity; with one moderately large tubercle each side of anterior area;lateral margins slightly narrowed centrally,distinctly widened at apex. Scutellum with yellowish-white pubescence not obscuring integument. Elytra: Humeri rounded, somewhat projected forward; with elevated centrobasal crest between humeri and scutellum, nearly conically elevat- ed anteriorly, covered with small tubercles, and distinct carina from apex of centrobasal crest to near apex; with another carina between the crest and humeri; elytral carina fused at their apex; coarsely, sparsely punctate, punctures sparser, shallower toward apex, with basal punctures anteriorly bordered by small tubercle; apex truncate, slightly concave centrally; pubescence mostly yellowish-white, less so between centrobasal crests, except wide, irregular, slightly oblique dark-brown pubescent band about middle (with irregular yellowish-white macula on inclined area), and dark-brown spots surrounding punctures; U-shaped white pubescent band on basal third partially lost in the specimen examined. Legs: Femora with yellowish-white pubescence partially obscuring integument on some areas, with brown pubescent areas interspersed on club. Tibiae with three rings of yellowish-white pubescence, one basally, one centrally, another at apex; remaining surface with brown pubescence. Protarsi with dark-brown pubescence dorsally, with a few whitish setae interspersed on tarsomere I. Tarsomere I of meso- and metatarsi with dense white pubescence on posterior ⅔ of dorsal surface, dark-brown pubescence basally; tarsomeres II-IV with dark-brown pubescence; tarsomere V with white pubescence not obscuring integument.

Abdomen: Ventrites with yellowish-white pubescence not obscuring integument, sparser centrally, with distal area of ventrites I-IV glabrous; distal margin of ventrite V concave.

Dimensions (mm): Total length, 12.80; prothoracic length, 2.60; anterior prothoracic width, 3.20; posterior prothoracic width, 3.15; maximum prothoracic width, 4.55; humeral width, 5.15; elytral length, 8.20.

Material examined: NICARAGUA (new country record): Nueva Segovia: Cerro Jesus (13°58′N, 86°10′W; 1,100-1,200 m), 1 male,VI-VII.2016, E. van den Berghe col. ( ACMT) .

Known geographical distribution ( Monné, 2019; Tavakilian & Chevillotte, 2019; new record): Guatemala, Honduras, Nicaragua.

Remarks: Bates (1880) described the species from Guatemala, based on a single female. Subsequently, the species has only been mentioned in catalogs and checklists, except for Turnbow et al. (2003), who provid- ed Honduras as a new country record. Examination of the photograph of the holotype, as well as the male described here, reveals that the eyes are finely granulated, and the elytra distinctly carinate. Accordingly, the species is transferred to Scythropopsis .

Figures

Figures 55-66. (55) Psapharochrus sallei, holotype female, dorsal habitus. (56‑61) Scythropopsis cornuta. (56) Holotype female, dorsal habitus. (57) Dorsal habitus, male. (58) Ventral habitus, male. (59) Lateral habitus, male. (60) Head, frontal view, male. (61) Lower eye lobe, male. (62‑66) Scythropopsis wappesi, male. (62) Dorsal habitus.(63)Ventral habitus.(64) Lateral habitus.(65) Head,frontal view.(66) Lower eye lobe.