Ismarus halidayi Förster, 1850

Kim, Chang-Jun, Notton, David G., Ødegaard, Frode & Lee, Jong-Wook, 2018, Review of the Palaearctic species of Ismaridae Thomson, 1858 (Hymenoptera: Diaprioidea), European Journal of Taxonomy 417, pp. 1-38 : 21-25

Ismarus halidayi Förster, 1850

Figs 1C, H , 2E

? Entomia campanulata Herrich-Schäffer, 1840: 127 (nomen dubium). Ismarus halidayi Förster, 1850: 285 .

Ismarus longicornis Thomson, 1858: 378 . Synonymized by Kolyada & Chemyreva (2016). Ismarus mongolicus Szabó, 1974: 23 . Synonymized by Kolyada & Chemyreva (2016).

Diagnosis

Antenna colour variable; POL as long as OOL in both sexes; A3 shorter than A 4 in both sexes; radial cell as long as length of marginal vein in both sexes; base of second tergite with long median furrow, at least to half of segment in both sexes.

Type material

MONGOLIA: ♀, holotype of I. mongolicus , Central aimak, Tosgoni ovoo, 5̅ 10 km N of Ulan-Baator, 1500̅ 1700 m, Exp. Dr. Z. Kaszab, 19̅20, 23̅ 24 Jul. 1968 ( HNHM, Typ. No. 2622, Mus. Budapest) holotype by original designation.

SWEDEN: ♀, lectotype of I. longicornis , “Sbg” [Sövdeborg], “Rh” [Carl David Emmanuel Roth leg.], “NHRS-HEVA 000003605” (NHRS) lectotype designated by Kolyada & Chemyreva (2016).

Additional material examined (91 ♀♀, 193 ♁♁)

CANADA: 1 ♀, Newfoundland , South Branch, Jul. 1974, MT, Heinrich leg. ( NHMUK010265151 ).

REPUBLIC OF IRELAND: 1 ♀, Co. Wicklow, Buckroney , 15 Jun. 1949, M.W.R. de V. Graham leg. ( NHMUK010265375 ).

GERMANY: 1 ♀, North Rhine-Westphalia, Leverkusen, Bergisch-Neukirchen, River Wupper , reared from flood debris, 3 Feb. 2002, M. Boness leg. ( DNPC); 1 ♁, same collecting data, 2 Jan. 1982 ( DNPC).

NORWAY: 1 ♀, Vestfold, Stokke, Melsomvik, 19 Jun. 2012, A. Staverløkk leg. (NINA); 1 ♀, Vestfold, Sandefjord, 20 Jun.2012,A.Staverløkk leg.(NINA);1♁,Vestfold, Horten, Løvøya, 27 May ̅ 16Jun.2014, MT, A. Staverløkk leg. (NINA); 1 ♁, Vestfold, Stokke, Feen, 19 Jun. 2012, A. Staverløkk leg. (NINA); 5 ♀♀, Aust-Agder, Froland, Øyrekjerr, 2 July-14 Aug. 2012, MT, A.Endrestøl (NINA); 1 ♀, same collecting data, 5 Jun.–2 Jul. 2015; 1 ♁, 1 ♀, Sogn og Fjordane, Sogndal, Fiksneset, 5 Jul. ̅ 10 Aug. 2011, MT, F. Ødegaard leg. (NINA); 1 ♀, Sør-Trøndelag, Oppdal, Helvetesdalen, 29 Jun.–17 Jul. 2014, MT, O. Hanssen leg. (NINA); 1 ♁, Sør-Trøndelag, Trondheim, Jonsvannet, Tangen, 31 May ̅ 30 Jun. 2015, MT, F. Ødegaard leg. (NINA); 2 ♀♀, Finnmark, Karasjok, Buddasnjarga, 69,371057° N, 25,815782° E, 26 Jun. ̅ 12 Jul. 2016, MT, F. Ødegaard leg. (NINA); 1 ♀, same collecting data, 12 Jul. ̅ 11 Aug. 2016.

SOUTH KOREA: 1 ♁, Chungcheongbuk Province, Chungju-si, Suanbo-myeon, Samun-ri , Mt. Woraksan , 35°49′46″ N, 128°04′05″ E, 16 Jun. ̅ 17 Jul. 2013, MT, J.K. Choi leg. ( YNU); 1 ♁ , Chungcheongbuk Province, Boeun-gun, Mt. Songni National Park, Beopjusamaepyoso , 36°32′06″ N, 127°49′40″ E, 12 May ̅ 21 Jun. 2007, MT, J.W. Lee leg. ( YNU); 1 ♁ , Chungcheongbuk Province, Boeun-gun Mt. Songnisan, Beopjusa, 36°32′06″ N, 127°49′40″ E, 5 May ̅ 31 Aug. 2011, MT, J.C. Jung leg. ( YNU); 4 ♁♁ , Chungcheongnam Province, Seosan-si, Haemi-myeon, Daegok-ri 880, Hanseo- Univ. , 36°41′30″ N, 126°34′50″ E, 14 May ̅ 11 Jun. 2009, MT, J.W. Lee leg. ( YNU); 1 ♁, Daejeon-si, Dong-gu, Daejeon-Univ., 13̅ 27 May 2006, MT, J.W. Lee leg. ( YNU); 9 ♁♁ , 1 ♀, same collecting data, 16 May ̅ 5 Jun. 2006, MT, J.W. Lee leg. (YNU); 2 ♁♁, 1 ♀, same collecting data, 12̅ 27 May 2007, MT, J.W. Lee leg. (YNU); 1 ♁, 6 ♀♀, Daejeon-si, Seo-gu, Polpyeong-dong, Wolpyeong Park, 36°20′30″ N, 127°21′30″ E, 12 May ̅ 24 Jun. 2008, MT, J.W. Lee leg. (YNU); 3 ♁♁, Daejon-si, Wadong, 36°24′02″ N, 127°25′98″ E, 6̅ 28 May 2006, MT, edge of wild rose patch, P. Tripotin leg. (CNCI); 10 ♁♁, 3 ♀♀, same collecting data, 28 May ̅ 19 Jun. 2006, MT, edge of wild rose patch, P. Tripotin leg. (CNCI); 1 ♀, same collecting data, 19 Jun. ̅ 16 Jul. 2006, MT, edge of wild rose patch, P. Tripotin leg. (CNCI); 12 ♁♁, 1 ♀, Gangwon Province, Chuncheon-si, Nam-myeon, Hudong-li , 25 May ̅ 14 Jun. 2003, MT, semishade, forest edge, P. Tripotin leg. ( CNCI); 1 ♁ , 2 ♀♀, Gangwon Province, Chuncheon, Nam-myeon, Hudong-li , 14 Jun. ̅ 6 Jul. 2003, MT, pasture, trail close to forest dege, P. Tripotin leg. ( CNCI); 7 ♁♁ , Gangwon Province, Chuncheon-si, Nam-myeon, Balsan , 300 m, 37°43′29″ N, 127°37′73″ E, 17 May ̅ 6 Jun. 2006, MT, forest, P. Tripotin leg. ( CNCI); 2 ♁♁, same collecting data, 26 Jun. ̅ 30 Jul. 2006, MT, forest, P. Tripotin leg. ( CNCI); 1 ♁ , Gangwon Province, Inje-gun, Girin-myeon, Mt. Bangtaesan , 37°53′41″ N, 128°21′21″ E, 24 Jun. ̅ 19 Jul. 2013, MT, J.W. Lee leg. ( YNU); 1 ♁ , Gangwon Province, Gwanggwon Prov. Environment Park, Bukbang, Hongchoen , 35°45′15.6″ N, 127°51′1.7″ E, 30 May ̅ 15 Jun. 2012, MT, S.J. Jang leg. ( YNU); 1 ♁ , Gangwon Province, Hoengseong-gun, Gapcheon-myeon, Hadae-ri , 37°31′34.14″ N, 128°09′05.03″ E, 26 May ̅ 2 Jun. 2009, MT, K.W. Lee leg. ( YNU) ; 1 ♀, Gangwon Province, Wonju-si, Heungeop-myeon, Yeonsei Univ. , Wonju Campus , 37°16′24″ N, 127°54′02″ E, 24 May ̅ 20 Jun. 2007, MT, H.Y. Han leg. ( YNU); 2 ♁♁ , 1 ♀, Gangwon Province, Wonjusi, Baegun-myeon, Mt. Baegunsan , 37°15′02″ N, 128°02′31″ E, 26 Jun. ̅ 15 Jul. 2007, MT, J.W. Lee leg. ( YNU); 4 ♁♁ , Gangwon Province, Wonju-si, Baegun-myeon, Mt. Baegunsan , 37°16′22″ N, 127°55′65″ E, 19 Jun. ̅ 5 Jul. 2011, MT, J.W. Lee leg. ( YNU); 1 ♁ , Gangwon Province, Wonju-si, Heungeop-myeon, Yeonsei Univ. , Wonju Campus , 37°16′24″ N, 127°54′02″ E, 24 May ̅ 20 Jun. 2007, MT, H.Y. Han leg. ( YNU) ; 1 ♀, Gangwon Province, Wonju-si, Heungeop-myeon, Maeji-ri , Yonsei Univ. , 37°16′53″ N, 127°54′02″ E, 19 May ̅ 6 Jun. 2011, MT, J.W. Lee leg. ( YNU) ; 1 ♀, same collecting data, 29 May ̅ 5 Jul. 2015, MT, H.Y. Han leg. (YNU); 2 ♁♁, Gangwon Province, Wonju-si, Sochomyeon, Hakgong-ri , Mt. Chiak , 37°22′18″ N, 128°03′1.84″ E, 30 May ̅ 8 Jun. 2013, MT, J.W. Lee leg.

(YNU); 1 ♀, same collecting data, 9̅ 20 Jun. 2013, MT, J.W. Lee leg. ( YNU); 1 ♀, Gyeonggi Province, Anvang-si, Manan-gun, Kwanag Arb. , 9̅ 24 Jun. 2007, MT , J.O. Lim leg. ( YNU); 1 ♁, 1 ♀, Gyeonggi Province, Gapyeong-gun, Goseong-ri, Mt. Homyeongsan , alt. 168 m, 37°43′15″ N, 127°29′18.9″ E, 1 May ̅ 26 Jun. 2009, MT , J.O. Lim leg. (YNU); 1 ♀, same collecting data, 26 Jun. ̅ 16 Jul. 2009, MT, J.O. Lim leg. (YNU); 4 ♁♁, same collecting data, 27 May ̅ 10 Jun. 2009, MT, J.O. Lim leg. ( YNU); 10 ♁♁, 1 ♀, Gyeonggi Province, Gapyeong-gun, Goseong-ri, Mt. Homyeongsan , alt. 200 m, 37°43′16.3″ N, 127°29′23.4″ E, 27 May ̅ 10 Jun. 2009, MT , J.O. Lim leg. ( YNU); 2 ♁♁, Gyeonggi Province, Gapyeong-gun, Cheongpyeong-myeon, Goseong-ri , Mt. Homyeongsan , alt. 220 m, 37°43′16.3″ N, 127°29′23.4″ E, 11̅ 25 Jun. 2009, MT , J.O. Lim leg. ( YNU); 5 ♁♁, 1 ♀, Gyeonggi Province, Gwangju-si, Docheog-myeon, Mt. Taehwasan , alt. 219 m, 37°18′05″ N, 127°19′01″ E, 8 May ̅ 7 Jun. 2007, MT , J.O. Lim leg. (YNU); 1 ♁, same collecting data, 25 May ̅ 8 Jun. 2007, MT, J.O. Lim leg. (YNU); 5 ♁♁, 2 ♀♀, same collecting data, 9̅ 24 Jun. 2007, MT, J.O. Lim leg. ( YNU); 8 ♁♁, 7 ♀♀, Gyeonggi Province, Namyangju-si, Choan-myeon, Songchon-ri , Mt. Ungilsan , alt. 134 m, 37°34′43″ N, 127°18′37″ E. 27 May ̅ 10 Jun. 2009, MT , J.O. Lim leg. (YNU); 2 ♀♀, same collecting data, 11̅ 25 Jun. 2009, MT, J.O. Lim leg. (YNU); 1 ♁, same collecting data, 1̅ 26 May 2009, MT, J.O. Lim leg. ( YNU); 1 ♁, Gyeonggi Province, Suwon-si, Kwonseon-gu, Seodon-dong , Suwon Arb. , alt. 42 m, 37°15′38.3″ N, 126°59′01.1″ E, 20 May ̅ 10 Jun. 2009, MT , J.O. Lim leg. ( YNU); 1 ♁, Gyeonggi Province, Yangpyeong-gun, Yongmun-myeon, Yeonsu-ri , Mt. Yongmunsan , 37°31′49.5″ N, 127°34′18.8″ E, 11̅ 25 Jun. 2009, MT , J.O. Lim leg. ( YNU); 1 ♁, Gyeongsangbuk Province, Cheongdogun, Gakbuk-myeon, Namsan-ri , 35°58′ N, 128°47′ E, 14 May ̅ 24 Jun. 2012, MT , J.W. Lee leg. ( YNU); 3 ♁♁, Gyeongsangbuk Province, Yeongju-si, Punggi-eup, Jungyeong , 35°53′42.7″ N, 128°26′22.0″ E, 21 May ̅ 3 Jun. 2009, MT , C.J. Kim leg. (YNU); 5 ♁♁, same collecting data, 3̅ 12 Jun. 2009, MT, C.J. Kim leg. (YNU); 5 ♁♁, same collecting data, 12̅ 22 Jun. 2009, MT, C.J. Kim leg. (YNU); 1 ♀, same collecting data, 22 Jun. ̅ 3 Jul. 2009, MT, C.J. Kim leg. ( YNU); 2 ♀♀, Gyeongsangnam Province, Hamyang-gun, Macheon-myeon, Samjeong-ri , Mt. Jirisan , alt. 700 m, 35°20′55″ N, 127°38′21″ E, 15 Jun. ̅ 22 Jul. 2003, MT, big clearing on forest edge , P. Tripotin leg. ( CNCI); 1 ♁, Gyeongsangnam Province, Samcheong-gun, Samjang-myeon, Yupyeon-ri, Wangdeungjae, Mt. Jiri National Park , 35°23′8.81″ N, 127°45′44.11″ E, 16 Jun. ̅ 20 Sep. 2011, MT , J.C. Jeong leg. ( YNU); 1 ♁, Jeju Province, Bukjeju-gun, Norooreum, Gongseong-ri , 33°21′59″ N, 126°26′22″ E, 19 May 2003, MT , J.W. Lee leg. ( YNU); 1 ♁, Jeollanam Province, Jangeong-gun, Bukha-myeon, Mt. Naejangsan , Sajabong , 36°24′14.01″ N, 126°52′12.09″ E, 21 Jun. 2005, MT , K.B. Kim leg. ( YNU); 1 ♁, Jeollanam Province, Jangeong-gun, Bukha-myeon, Mt. Naejangsan , Baegyangsa , 35°26′22.8″ N, 126°53′00″ E, 18 Apr. ̅ 30 May 2008, MT , J.W. Lee leg. (YNU); 1 ♀, same collecting data, 30 May ̅ 5 Aug. 2008, MT, J.W. Lee leg. ( YNU); 1 ♁, Jeollabuk Province, Jeongeup-si, Ibam-myeon, Deungcheon-ri , Wetland 35°28′35.95″ N, 126°47′59.17″ E, 21 Jun. 2005, MT , M.B. Yun leg. ( YNU); 1 ♁, Jeollabuk Province, Jeongeup-si, Naejang-dong, Mt. Naejangsan , Wonjeogam , 35°29′36″ N, 126°53′37″ E, 13̅ 29 May 2007, MT , J.W. Lee leg. (YNU); 2 ♁♁, same collecting data, 20 Jun. 2005, MT, J.W. Lee leg. ( YNU); 1 ♀, Jeollabuk Province, Jeongeup-si, Singjeong-dong, Mt. Naejangsan , Namchanggol , 35°27′48″ N, 126°50′18″ E, 30 May ̅ 5 Jul 2008, MT , J.W. Lee leg. ( YNU); 8 ♁♁, 3 ♀♀, Jeollabuk Province, Jeongeup-si, Yongsan-dong , 35°19′8.97″ N, 126°53′11.74″ E, 19 May ̅ 19 Jun. 2004, MT , M.K. Yun leg. (YNU); 1 ♁, same collecting data, 4 Jun. 2006, MT, J.W. Lee leg. ( YNU); 10 ♁♁, Jeollanam Province, Jirisan Hamyang-gun, Macheon-myeon, Samjeong-li , 700 m, 35°20′55″ N, 127°38′21″ E, 15̅ 31 May 2003, MT, forest clearing, stream , P. Tripotin leg. ( CNCI); 1 ♁, Jeollanam Province, Jirisan, Hamyanggun, Macheon-myeon , Samjeong-li , 700 m, 35°20′55″ N, 127°38′21″ E, 1̅ 15 Jun. 2003, MT, big clearing, forest edge , P. Tripotin leg. ( CNCI); 1 ♁, Jeollanam Province, Jirisan, Hamyang-gun, Macheonmyeon , Samjeong-li , 700 m, 35°20′55″ N, 127°38′21″ E, 11̅ 18 May 2003, MT, in big clearing , P. Tripotin leg. ( CNCI); 1 ♁, Jeollanam Province, Jirisan, Piagol Valley, Jikjeok , 35°16′39″ N, 127°33′ E, 1̅ 15 Jun. 2003, MT, in shade in small clearing in forest , C. Young leg. ( CNCI); 1 ♁, 1 ♀, Jeollanam Province, Yeongam-gun, Hoemun-ri, Daedongjae , 22 May ̅ 31 Jul. 2010, MT , J.K. Kim leg. ( YNU); 9 ♁♁, Seoul-si, Jongno-gu, Gugi-dong , Gugi Valley , Mt. Bukhan National Park , 35°37′11″ N, 126°57′42″ E, 5 Jun. ̅ 22 Jul. 2010, MT , J.C. Jeong leg. ( YNU); 1 ♀, Gyeongsangbuk Province, Mungyeong-si, Gaeun-eup, Wonjang-ri, Mt. Songnisan, Beorimgijae , 36°40′59″ N, 127°57′07″ E, 21 May ̅ 15 Jun. 2013 , J.K. Choi leg. (YNU).

JAPAN: 1 ♀, Aichi, Mt. Chausu , alt. 1300 m (ssw). 9 Jul. 1995, K. Yamagishi leg. ( CNCI) ; 7 ♁♁, 1 ♀, Hokkaido, Sapporo , 24̅ 29 Jul. 1988, MT, K. Maeto leg. ( CNCI) ; 1 ♁, Hokkaido, Sapporo, Hitsujgaoka, Hokkaido , National Agriculture Experiment station , alt. 133 m, 43°00′30″ N, 141°24′47.9″ E, 30 Jun. ̅ 2 Jul. 2008, MT, J.W. Lee leg. ( YNU) ; 1 ♀, Nishioka-Park, Toyohira-ku, Sapporo-shi , Hokkaido, alt. 140 m, 42°59′19.3″ N, 141°22′46.9″ E, 27 Jul. 2013, MT, J.W. Lee leg. ( YNU) .

RUSSIA: 1 ♁, Primorsky-Krai , Vladivostok, 8̅ 15 Jun. 2008, MT, J.W. Lee leg. ( YNU); 1 ♀, Primorsky- Krai, Vladivostok, 43°15′29″ N, 132°02′12.71″ E, 15̅ 20 Jun. 2008, MT, J.W. Lee leg. ( YNU); 1 ♁, Primorsky-Krai, Vladivostok , 20 Jun. ̅ 5 Jul. 2008, MT, J.W. Lee leg. (YNU).

SWEDEN: 1 ♁, Västerbotten, Hällnäs , 3 Jul. 1960, K.- J. Hedqvist leg. ( NHMUK010265149 ) ; 2 ♁♁, Skåne, Ringsjön , 4 Jun. 1938, D.M.S. Perkins and J.F. Perkins leg. ( NHMUK010265147 , NHMUK010265148 ) ; 1 ♀, same collecting data, 24 Jun. 1938 (NHMUK010265143); 1 ♀, same collecting data, 27 Jun. 1938 (NHMUK010265144); 1 ♀, Uppland, Vallentuna , Jun. 1957, K.- J. Hedqvist leg. ( NHMUK010265146 ) ; 1 ♀, same collecting data, 31 May 2007 (NHMUK010265145).

UNITED KINGDOM: 1 ♀, England, Beds, Flitwick Moor, TL045350, Betula , 25 Jun. 1960, V.H. Chambers leg. (NHMUK010265076); 1 ♁, same collecting data (NHMUK010265092); 1 ♀, same collecting data, Lonicera / Betula / Quercus , 19 Jun. 1980 (NHMUK010265078); 1 ♁, same collecting data, Lonicera , 29 Jun. 1981 (NHMUK010265091); 1 ♀, England, Beds, Flitwick Moor, TL045350, 17 Jul. 1981, V.H. Chambers leg. (NHMUK010265080); 1 ♀, England, Beds, Kings Wood, SP920294, Betula , 11 Jul. 1951, V.H. Chambers leg. (NHMUK010265074); 1 ♁, same collecting data, Betula , 21 Jun. 1953 (NHMUK010265087); 1 ♀, same collecting data, Betula , 26 Jun. 1953 (NHMUK010265075); 1 ♀, same collecting data, Betula , 3 Aug. 1953 (NHMUK010265084); 1 ♁, same collecting data, bred, host collected 11 Jul. 1953, wasp emerged 16 Jun. 1954, ex Anteon sp. on Oncopsis sp. on Betula sp. (NHMUK010265089); 1 ♁, England, Beds, path to Sharpenhoe Clappers, TL063298, Alnus glutinosus , 12 Jul. 1951, V.H. Chambers leg. (NHMUK010265088); 1 ♀, England, Beds, Sutton Fen, TL202475, Betula , 18 Jul. 1978, V.H. Chambers leg. (NHMUK010265079); 1 ♀, same collecting data, fern, 2 Jul. 1980 (NHMUK010265077); 1 ♀, England, Bucks, Soulsbury, Rammamere Heath, 4̅ 17 Jul. 1944, R.B. Benson leg. (NHMUK010265072); 1 ♀, England, Bucks, Wootton Underwood, 4 Jul. 1957, R.B. Benson leg. (NHMUK010265081); 1 ♀, England, Cambs, Chippenham Fen, TL650693, carr / reedbed,?̅ 6 Jul. 1983, MT, J. Field leg. (DNPC); 1 ♀, England, Ches, Abbots Moss, SJ596680, by stream, 1̅ 19 Jun. 1990, MT, D.G. Notton leg. (DNPC); 1 ♁, same collecting data, 19 Jun. ̅ 10 Jul. 1990 (DNPC); 1 ♀, England, Devon, Dartmoor, Lustleigh, Jun. 1934, R.C.L. Perkins leg. (NHMUK010265071); 1 ♁, Scotland, Glasgow, Clober nr Milngavie, pre-1910, P. Cameron leg. (NHMUK010265139); 1 ♁, England, Hants, Farley, 12 Jun. 1938, R.B. Benson leg. (NHMUK010265095); 1 ♀, England, Herts, Tring, 16 Jul. 1942, R.B. Benson leg. (NHMUK010265073); 1 ♀, Scotland, Highland, Aviemore, 30 Jun. 1934, R.B. Benson leg. and J.E. Benson leg. (NHMUK010265138); 1 ♁, same collecting data, 4 Jun. 1952, R.B. Benson leg. (NHMUK010265142); 1 ♁, Scotland, Highland, Bonar, pre-1910, P. Cameron leg. (NHMUK010265086); 1 ♁, same collecting data, pre-1909 (NHMUK010265141); 1 ♀, Scotland, Highland, Kinlochewe, 1̅ 8 Jun. 1961, R.B. Benson leg. (NHMUK010265137); 1 ♀, England, Lancs, Freshfield, 28 Jul. 1961, M.W.R. de V. Graham leg. (NHMUK010265082); 1 ♀, England, Norfolk, Santon Downham, TL818883, MT,? ̅ 6 Jul. 1985, J. Field leg. (DNPC); 2 ♀♀, Scotland, Perth and Kinross, Ballinluig, 20 Jul. 1977, J.S. Noyes, L. Rogers and T. Huddleston leg. (NHMUK010265097, NHMUK010265136); 1 ♁, England, Suffolk, Brandon, 23̅ 30 May 1945, R.B. Benson leg. (NHMUK010265096); 1 ♀, England, Surrey, Horsley, 12 Jun. 1957, J.F. Perkins leg. (NHMUK010265083); 1 ♁, England, Surrey, Oxshot, 25 Jun. 1937, G.E.J. Nixon leg. (NHMUK010265094); 2 ♁♁, England, 12 Jun. 1966, V.H. Chambers leg. (NHMUK010265085, NHMUK010265090).

Variation

Body length 1.93̅ 3.79 mm in both sexes; antenna colour variable in both sexes: totally dark brown or brown, dark brown or brown except segments 1 or 1̅2 to 1̅10 yellowish; anterior scutellar pit with median keel present or absent, weakly crenulate or completely smooth at bottom in both sexes; hind leg colour variable in both sexes: femur yellow to brown, tibia and tarsus yellow to dark brown; median furrow extending 0.5̅0.9 × length of second tergite in both sexes.

Distribution

Japan (new record), South Korea (new record), Azerbaijan ( Kolyada & Chemyreva 2016), Bulgaria ( Petrov1990), Canada ( Masner 1976), China (Ningxia, Sichuan, Guizhou, Yunnan, Tibet) ( Liu et al. 2011), Czech Republic ( Hellén 1964), Denmark ( Johnson 2016), Finland ( Hellén 1964), Georgia ( Kolyada & Chemyreva 2016), Germany ( Kieffer 1916), Hungary ( Kieffer 1916; Peeters 2015), Mongolia ( Szabó 1974), Netherlands ( Peeters 2015), Norway (http://www.biodiversity.no/Pages/135494), Republic of Ireland ( Stelfox 1966; O’Connor et al. 2004), Russia (European, Far East, Siberia) ( Kolyada & Chemyreva 2016), Scotland ( Nixon 1957), Sweden ( Nixon 1957), United Kingdom ( Kieffer 1916; Nixon 1957, Notton 1996; O’Connor et al. 2004), USA ( Masner 1976).

Host

Reared from Anteon jurineanum Latreille, 1809 ( Chambers 1955, as A. brevicorne ; Olmi 2000) and A. infectum (Haliday) (Dryinidae) ( Chambers 1955, 1981), both in United Kingdom. However, Chambers (1981: as I. halidayi ), but not Chambers (1955), which is in fact I. similis sp. nov., see below.

Notes

Kolyada & Chemyreva (2016) incorrectly state that Entomia campanulata is a nomen oblitum, however no action has been taken under Article 23.9.2 ( ICZN 1999) to reverse the precedence with respect to I. halidayi so it cannot be a nomen oblitum. We follow Masner (1976) who doubtfully included in it synonymy with Ismarus halidayi and we consider that the interpretation of the species is doubtful and so the synonymy is also doubtful, and it is therefore a nomen dubium, i.e., although it is the older name, it is not certainly the valid name for the species referred to here as I. halidayi .

Figures

Fig. 1. Ismarus spp., ♀♀ (A–G. Habitus; H–I. Mesosoma in lateral view). A. I. apicalis Kolyada & Chemyreva, 2016.B. I.dorsiger (Haliday, 1831).C,H. I. halidayiFörster, 1850.D.I. grandisAlekseev, 1978. E. I. rugulosus Förster, 1850. F. I. spinalis Kolyada & Chemyreva, 2016. G, I. I. flavicornis (Thomson, 1858).

Fig. 2. Ismarus spp., ♁♁. (A–G, I. Habitus; H. Antennae). A. I. apicalis Kolyada & Chemyreva, 2016. B. I. dorsiger (Haliday, 1831). C. I. flavicornis (Thomson, 1858). D. I. grandisAlekseev, 1978. E. I. halidayi Förster, 1850. F. I. multiporus Kolyada & Chemyreva. G. I. rugulosus Förster, 1850. H–I. I. spinalis Kolyada & Chemyreva, 2016.