Crassignatha nasalis Y. Lin & S. Li, 2020

Li, Ya, Lin, Yucheng & Li, Shuqiang, 2020, A review of Crassignatha (Araneae, Symphytognathidae), ZooKeys 988, pp. 63-128 : 63

publication ID	https://dx.doi.org/10.3897/zookeys.988.56188
publication LSID	lsid:zoobank.org:pub:6E64D69B-DD73-4A7E-AE2B-3CD21247A5E3
persistent identifier	https://treatment.plazi.org/id/7084BD59-EC9E-44F1-9DC2-F0E08117E7A7
taxon LSID	lsid:zoobank.org:act:7084BD59-EC9E-44F1-9DC2-F0E08117E7A7
treatment provided by	ZooKeys by Pensoft
scientific name	Crassignatha nasalis Y. Lin & S. Li
status	sp. nov.

Treatment

Crassignatha nasalis Y. Lin & S. Li sp. nov. Figs 16 View Figure 16 , 17 View Figure 17 , 38 View Figure 38

Typ e material.

Holotype ♂ (NHMSU Ar 026) and paratypes 9♀ (NHMSU Ar 027-035), China: Sichuan Province, Luzhou City, Gulin County, Yuhua Township, Taoyuan Cave (27.98293°N, 105.99833°E; 910 m), 21.IV.2014, Y. Lin, H. Zhao, and Y. Li leg. 1♂ (NHMSU-HA041) and 1♀ (NHMSU-HA041) used for sequencing, GenBank: MT991992 and MT991991, same data as for preceding.

Diagnosis.

The male of C. nasalis sp. nov. is similar to that of C. rostriformis sp. nov. but can be distinguished by the sharp hook of the median apophysis and a straight cymbial tooth vs. a blunt hook of the median apophysis and a hook-like cymbial tooth (Figs 17A, B View Figure 17 , 25A, B View Figure 25 ). The female is similar to C. quanqu in epigyne shape but differs from the latter by the indistinct scape and the copulatory ducts nearly forming a closed rhombic area at center of vulva vs. obvious scape and copulatory ducts not forming a closed area at center of vulva (Fig. 17E, F View Figure 17 vs. Fig. 23D, E View Figure 23 ).

Description.

Male (holotype). Total length 0.80. Carapace 0.40 long, 0.32 wide, 0.44 high. Clypeus 0.12 high. Sternum 0.24 long, 0.20 wide. Abdomen 0.52 long, 0.44 wide, 0.56 high. Length of legs: I 1.10 (0.30, 0.14, 0.30, 0.16, 0.20); II 0.92 (0.24, 0.12, 0.24, 0.14, 0.18); III 0.72 (0.20, 0.10, 0.14, 0.10, 0.18); IV 0.82 (0.22, 0.10, 0.20, 0.12, 0.18).

Somatic characters (Fig. 16A-C View Figure 16 ). Coloration: carapace, sternum, chelicerae, endites, and labium brown. Legs yellow-brown. Abdomen pale gray, ventrally darker than dorsally, with sclerotized dots and sparse setae; posterolateral scutum light brown. Prosoma: carapace nearly rounded, thoracic center smooth, cephalic area and margins granulated and pitted, with two strong setae medially. Cephalic area strongly elevated. ALE protruded, PER strongly recurved. Clypeus concave. Chelicerae covered with setae anteriorly. Sternum almost heart shaped, slightly plump, truncated posteriorly, surface rough. Legs: tibia II with two large clasping spurs. Abdomen: sub-rounded dorsally. Spinnerets weakly sclerotized, with a circular plate.

Palp (Fig. 17A-C View Figure 17 ): pale, weakly sclerotized. Cymbium with a few setae apically, cymbial tooth spur-like, located at distal terminus. Tegulum globose, smooth. Plate-like median apophysis with a spike-shaped distal process. Embolic membrane arises behind the median apophysis. Embolus short, rigid, basally constricted and mesally widened, distally forming an inverted Z-shape.

Female (one of paratypes). Total length 1.00. Carapace 0.40 long, 0.36 wide, 0.40 high. Clypeus 0.16 high. Sternum 0.24 long, 0.24 wide. Abdomen 0.68 long, 0.68 wide, 0.76 high. Length of legs: I 1.40 (0.46, 0.16, 0.32, 0.20, 0.26); II 1.18 (0.36, 0.14, 0.26, 0.18, 0.24); III 0.82 (0.20, 0.12, 0.16, 0.14, 0.20); IV 1.04 (0.30, 0.14, 0.24, 0.16, 0.20).

Somatic characters (Fig. 16D-F View Figure 16 ). Coloration: prosoma and legs as in male. Abdomen dark, with tiny light yellow dots. Prosoma: carapace nearly pear shaped, ocular arrangement and modification as in male. Cephalic area lower than in male. PER straight. Mouthparts and sternum as in male. Abdomen: globular dorsally. Spinnerets weakly sclerotized.

Epigyne (Fig. 17D-F View Figure 17 ): epigynal area distinctly sclerotized, with a few setae. Scape short, strongly sclerotized. Copulatory openings separated, resembling a pig snout. Vulva visible via translucent tegument. Paired spermathecae separated by less than ½ their diameter. Fertilization ducts originate inside posterior edge of spermathecae, curving toward center of spermathecae. Copulatory ducts connected to dorsal center surface of spermathecae, twisting 3 ×, then forming a nearly closed rhombic area in center of vulva.

Etymology.

The specific epithet is a Latin adjective (= nasal) and refers to the shape of the copulatory openings of the epigyne.

Distribution.

China (Sichuan) (Fig. 38 View Figure 38 ).