Anastrophyllum nigrescens (Mitt.) Steph., Hedwigia

Váňa, Jiří, Söderström, Lars, Hagborg, Anders & von Konrat, Matt, 2013, Notes on Early Land Plants Today. 33. Notes on Anastrophyllaceae (Marchantiophyta), Phytotaxa 81 (1), pp. 26-32 : 28-30

Anastrophyllum nigrescens (Mitt.) Steph., Hedwigia 32: 140, 1893 ( Stephani 1893).

Basionym:— Jungermannia nigrescens Mitt. , Hooker's J. Bot. Kew Gard. Misc. 3: 358, 1851 ( Mitten 1851).

= Anastrophyllum paramicola R.M.Schust., Beih. Nova Hedwigia 119: 316, 2002 ( Schuster 2002a), syn. nov. Type :— ECUADOR. Hyperpáramo , crest of road from Pifo to Papallacta, R.M. Schuster RMS 93-220 (holotype F, non vidi).

Anastrophyllum auritum (Lehm.) Steph., Bull. Herb. Boissier ser. 2 1: 1137 (Spec. Hepat. [Stephani] 2: 120), 1901 ( Stephani 1901).

Basionym:— Jungermannia aurita Lehm. , Linnaea 4: 368, 1829 ( Lehmann 1829).

Type : — SOUTH AFRICA. Table Mt., Ecklon (holotype S!, isotypes H-SOL!, W-Lindenb. Hep. 250!) .

Ξ Sarcocyphos auritus (Lehm.) Nees, Syn. Hepat. 1: 9, 1844 ( Gottsche et al. 1844) Ξ Nardia aurita (Lehm.) Trevis., Mem. Reale Ist. Lombardo Sci., Ser. 3, C. Sci. Mat. 4: 401, 1877 ( Trevisan 1877) Ξ Marsupella aurita (Lehm.) Sim, Trans. Roy. Soc. South Africa 15: 112, 1926 ( Sim 1926). = Jungermannia crebrifolia Hook. f. et Taylor, London J. Bot. 3: 467, 1844 ( Hooker & Taylor 1844), syn. fide Váňa in

Gradstein & Costa (2003). Lectotype:— CHILE. Prov. Magallanes: Cabo de Hornos (= Cape Horn), 1843, Hooker

(Voyage of H. M. S. Erebus & Terror) (FH!, isolectotypes BM!, G-121797! [=G-17223!] 1, NY!, S!, W-Lindenb.

Hep. 1627!) Ξ Anastrophyllum crebrifolium (Hook.f. et Taylor) Steph., Hedwigia 32: 140, 1893 ( Stephani 1893).

1. Citation of specimens in G should preferably use the barcode (M. Price, pers. comm.) but for comparability the numbers printed on the specimen, which have often been cited by previous authors, are also given here in square brackets. In case the barcode number is missing, the older number is still given within square brackets.

= Jungermannia leucocephala Taylor ex Lehm. , Nov. Stirp. Pug. 8: 8, 1844 ( Lehmann 1844), syn. of A. crebrifolium fide Engel (1978). Lectotype:— ECUADOR. apud Cayambe, 14.000 ft., 1827, Jameson (FH!, isolectotypes BM!, NY) Ξ Anastrophyllum leucocephalum (Taylor ex Lehm.) Steph., Hedwigia 32: 140, 1893 ( Stephani 1893).

= Jungermannia cryptodon Wilson ex Hook. f. et Taylor, London J. Bot. 3: 467, 1844 ( Hooker & Taylor 1844); nom. inval. (Art. 32.1.d; no description), Bot. Antarctic Voy. 1, Fl. Antarct. 2: 434, 1847 ( Hooker 1847), syn. nov. Original material:— COLOMBIA. Pilzhum, 25 August 1842, Mathews (FH!).

= Jungermannia leucostoma Taylor, London J. Bot. 5: 272, 1846 ( Taylor 1846), syn. fide Váňa (1984). Type:— PERU. Jameson (holotype FH!). Ξ Anastrophyllum leucostomum (Taylor) Steph., Hedwigia 32: 140, 1893 ( Stephani 1893).

= Jungermannia intricata Lindenb. et Gottsche, Syn. Hepat. 5: 679, 1847 ( Gottsche et al. 1847), syn. nov. Lectotype:— MEXICO. prope Yavesea, June 1842, Liebman 488b (C-Pl. mexic. 10348!, isolectotype [G-16447!]) Ξ Sphenolobus intricatus (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier ser. 2, 2: 167 (Spec. Hepat. [Stephani] 2: 159), 1902 ( Stephani 1902b) Ξ Eremonotus intricatus (Lindenb. et Gottsche) R.M.Schust., Rev. Bryol. Lichénol. 34: 269, 1966 ( Schuster 1966a) Ξ Anastrophyllum intricatum (Lindenb. et Gottsche) R.M.Schust., Hepat. Anthocerotae N. America 2: 709, 1969 ( Schuster 1969).

= Jungermannia lechleri Gottsche et Hampe , Linnaea 27 (= n. ser. vol. 11): 554, 1854 [1856] ( Hampe 1854), syn. nov. Lectotype:— PERU. St. Gavan, July 1854, Lechler (Plantae Peruvianae 2254) (BM!, isolectotypes G-121846! [=G- 17264!], G-121848 [=G-17273!], H-SOL!, YU!, W!) Ξ Anastrophyllum lechleri (Gottsche et Hampe) Steph., Hedwigia 32: 140, 1893 ( Stephani 1893).

= Jungermannia calocysta Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 517, 1885 ( Spruce 1885), syn. fide Váňa (1984). Type:— ECUADOR. Andes Quitenses, prope Loja, Massee (non vidi) Ξ Anastrophyllum calocystum (Spruce) Steph., Hedwigia 32: 140, 1893 ( Stephani 1893).

= Anastrophyllum apertifolium Steph., Bull. Herb. Boissier ser. 2, 1: 1134 (Spec. Hepat. [Stephani] 2: 117), 1901. ( Stephani 1901), syn. nov. Type :— PERU. St. Gavan, Lechler (holotype G-66728 [=G-11065]).

= Anastrophyllum glaziovii Steph., Bull. Herb. Boissier ser. 2, 1: 1138 (Spec. Hepat. [Stephani] 2: 121), 1901 ( Stephani 1901), syn. fide Váňa in Gradstein & Costa (2003). Type:— BRAZIL. Rio de Janeiro: Glaziou 4534 (lectotype G- 66708! [=G-11163!], isolectotypes BM!, FH!, G-60705, S!).

= Anastrophyllum mandonii Steph., Bull. Herb. Boissier ser. 2, 1: 1139 (Spec. Hepat. [Stephani] 2: 122), 1901 ( Stephani 1901), syn. nov. Type :— BOLIVIA. Mandon 808 p. p. (holotype G-66727! [=G-17263!]).

= Anastrophyllum capillaceum Steph., Arch. Mus. Nac. Rio de Janeiro 13: 110, 1903 ( Stephani 1903), syn. fide Váňa in Gradstein & Costa (2003). Type:— BRAZIL. Serra do Itatiaya, 2100 m, 13. June 1902, Dusén 365 (holotype G- 66740! [=G17214!]).

= Anastrophyllum gambaragarae Gola, Ann. Bot. (Rome) . 6: 274, 1907 ( Gola 1907), syn. fide Váňa (1975). Type:— UGANDA. Ruwenzori, Bujongolo valle Mobuku, 3800 m, Duke of Apruti Exped. (FT!).

= Anastrophyllum calcaratum Steph., Wiss. Ergebn. Deut. Zentr. -Afr. Exped. (1907-08), Bot. 2: 113, 1910 ( Mildbraed 1910), syn. fide Váňa (1975). Lectotype ( Váňa 1975):—Africa, Kilimanjaro Mt, 1901, Uhlig (G-45023! [=G- 16418!]).

= Anastrophyllum grossitextum Steph., Wiss. Ergebn. Deut. Zentr. -Afr. Exped. (1907-08), Bot. 2: 114, 1910 ( Mildbraed 1910), syn. fide Váňa (1975). Type:— UGANDA. Ruwenzori, Butagu Valley, 3300 m, Mildbraed 2621 ex p. (holotype G-45024 [=G-16428!]).

= Anastrophyllum semifissum Steph., Kungl. Svenska Vetenskapsakad. Handl. 46 (9): 21, 1911 ( Stephani 1911) syn. of A. crebrifolium fide Engel (1978). Lectotype ( Engel 1978):— CHILE. Prov. Magallanes: Canal Gajardo, Cta. Inga, 27 April 1908, Halle et Skottsberg 626 (UPS!, isolectotypes in BM!, S!).

= Sphenolobus savioi Gola, Mem. Reale Accad. Sci. Torino, Ser. 2, 65: 4, 1914 [1916] ( Gola 1916), syn. fide Váňa (1975). Type:— KENYA. Mt. Kinangop prope M. Kenya, 2000 m, Savio (holotype TOM).

= Anastrophyllum cuspidatum Steph., Biblioth. Bot. 87: 186, 1916 ( Stephani 1916), syn. nov. Type :— BOLIVIA. Tablas, 1912, Herzog 2907 (holotype G-66729 [=G-11068!]).

[= Anastrophyllum hians Steph., Biblioth. Bot. 87: 186, 1916 p.p. ( Stephani 1916), p. p., syn. nov. Type:— BOLIVIA. San Mateo , Sunchal, leg. Herzog 4473 (paratype G-282089! [=G-18864!]). Note:—The lectotype, Herzog 5077, is Anastrophyllum piligerum (Nees) Steph. ]

= Marsupella pusilla Steph., Biblioth. Bot. 87: 181, 1916 ( Stephani 1916), syn. fide Gradstein et al. (2003). Type:— BOLIVIA. Cerro Tunari, 4700 m, Herzog 4891 (holotype G-67517! [=G-14599!]).

= Anastrophyllum parvum Steph., Biblioth. Bot. 87: 187, 1916 ( Stephani 1916), nom. inval. (Art. 32,1,d; no description), syn. nov. Original material:— BOLIVIA. Waldgrenze über Tablas, 3400 m, May 1911, Herzog 2881 (JE!, W!).

= Anastrophyllum hintzeanum Steph., Sp. Hepat. (Stephani) 6: 106, 1917 ( Stephani 1917), syn. fide Váňa (1975). Type:— CAMEROON. 4000 m, Hintze 26d (holotype G-45025 [=G-16429!]).

= Anastrophyllum inaequale Steph., Sp. Hepat. (Stephani) 6: 106, 1917 ( Stephani 1917), syn. nov. Type:— BOLIVIA. Prov. Cochabamba: “las Estradillas”, prope Incacorral , 3000-3200 m, January 1908, Herzog (herb. Levier 6027) (holotype G-66731! [=G-17240!], isotypes FI!, YU!).

= Marsupella crenulata C.Massal. et Steph., Atti Reale Ist. Veneto Sci. Lett. Arti. 87: 236, 1928 ( Massalongo 1928), nom. illeg. (Art. 53.1; hom. illeg. [non Spruce 1881]), syn. fide Váňa et al. (2010). Type:— ARGENTINA. Tierra del Fuego: ad tundras alpinas e Staten Island, 2. March 1928, Spegazzini (holotype VER).

= Anastrophyllum minutirete Herzog, Hedwigia 74: 85, 1935 ( Herzog 1935), syn. nov. Lectotype:— COLOMBIA. Páramo El Boquerón bei Bogota, 3200 m, 1929, Troll 2155 (JE!, isolectotype S!).

= Anastrophyllum microdictyon Herzog, Beih. Bot. Centralbl. 61B: 563, 1942 ( Herzog 1942), syn. nov. Lectotype:— BOLIVIA. zwischen San Mateo und Sunchal, 2800 m, April 1911, Herzog 4449 (JE-H-1536!).

= Anastrophyllum leucostomum var. capillaceum Schiffn., Österr. Akad. Wiss., Math. -Naturwiss. Kl., Denkschr. 111: 42, 1964 ( Schiffner & Arnell 1964), syn. fide Váňa in Gradstein & Costa (2003). Type:— BRAZIL. ad confines Rio de Janeiro –Minas Geraës, in rupestribus montis Itatiaya, locis umbrosis humidis, 1300-2750 m, 18. September 1901, Schiffner 437 (holotype W!).

= Anastrophyllum gottscheanum R.M.Schust., Rev. Bryol. Lichénol. 34: 270, 1966 ( Schuster 1966a), syn. nov. Type :— MEXICO. Pico de Orizaba, Liebman (ex Herb. Gottsche) (holotype G-113582* [=G-17238!]).

= Anastrophyllum novazelandiae R.M.Schust., Rev. Bryol. Lichénol. 34: 282, 1966 ( Schuster 1966a). syn. nov. (ex description, illustration and specimens seen in F). Type :— NEW ZEALAND. South I.: Fiordland Nat. Park , Route Burn track between Earland Falls and Lake Mackenzie, W slope of Humboldt Mts., Upper Hollyford Valley, 4200- 4500 ft., Schuster 49471b (holotype F, non vidi).

= Anastrophyllum crenulatum R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 740, 1969 ( Schuster 1969), syn. nov. (ex description and illustration). Type:— ARGENTINA. Tierra del Fuego: Cerro Garibaldi, Schuster 58304a (holotype F, non vidi).

= Anastrophyllum appendiculatum N.Kitag., J. Hattori Bot. Lab. 33: 210, 1970 ( Kitagawa 1970), syn. fide Váňa (1975). Type:— MALAYSIA. Sabah: Mt. Kinabalu, granodiorite slope between Low´s Peak and Sayat Sayat, 3700-4050 m, 1963, Mizutani 2912 (holotype NICH-252912!).

= Anastrophyllum hypocladopsis R.M.Schust., Beih. Nova Hedwigia 119: 315, 2002 ( Schuster 2002a), syn. nov. (ex description and illustration). Type:— VENEZUELA. Estado Tachira: Páramo de Tama , Villa Paez , 3000 m, Schuster 76-1941a (holotype F, non vidi).

= Anastrophyllum intermedium R.M.Schust., Beih. Nova Hedwigia 119: 303, 2002 ( Schuster 2002a), nom. inval. (Art. 37.7; herbarium not indicated), syn. nov. (ex description and illustration). Original material:— VENEZUELA. Estado Korida: Sierra Nevada de Korida, above Loma Redonda, Schuster 76-1411c (F, non vidi).

= Anastrophyllum ambiguum R.M.Schust., Beih. Nova Hedwigia 119: 305, 2002 ( Schuster 2002a), nom. inval. (Art. 36.1; no Latin description), syn. nov. (ex description). Original material:— NEW ZEALAND.