Xantholinus (Typhlolinus) reitteri Coiffait, 1966

Assing, Volker, 2007, On the Xantholinini of Turkey and adjacent regions (Coleoptera: Staphylinidae: Staphylininae), Zootaxa 1474, pp. 1-54 : 17-18

Xantholinus (Typhlolinus) reitteri Coiffait, 1966 a

( Figs. 22–26 , 98 )

Xantholinus (Acanthophallus) reitteri Coiffait, 1966a: 198 f.

Xantholinus (Acanthophallus) svanetskiensis Coiffait, 1973: 108 f.; syn. n. Xantholinus (Helicophallus) motschulskyi Bordoni, 1999a: 4 ; syn. n.

Type material examined. X. reitteri : Holotype: ɗ, "Caucasus. Swanetien. Leder. Reitter / Xanth. fortepunctatus Mots. Coll. Reitter / Type / Xantholinus reitteri Coiff. H. Coiffait det. 1964 / Xantholinus reitteri Coiffait det. V. Assing 2007 " ( HNHM).

X. svanetskiensis : Holotype: ɗ, "Svania / Brotherus / Holotypus / Xantholinus (Acanthophallus) svanetskiensis H. Coiffait 1973 / Xantholinus reitteri Coiffait det. V. Assing 2007 " (MNHNP).

X. motschulskyi : Lectotype, present designation: ɗ, "Gilan ( Iran), Heinz leg. / Masuleh, 1.VIII.67, südl. Talysch, 1000–1800 m / Lectotypus ɗ Xantholinus motschulskyi Bordoni , desig. V. Assing 2006 / Xantholinus reitteri Coiffait det. V. Assing, 2006" (cAss). Paralectotypes: 1ɗ, Gilan ( Iran), Heinz leg. / Javaherdeh, 16.VII.75, südl. Ramsar, 1800–2000 m (cKor); 1ɗ, Naharkhoran, 24.VIII.67, südl. Gorgan, 450–1000 m, Heinz leg. (cKor). Additional unexamined paralectotypes are deposited in cKor: according to Korge (1973) 9 exs. with the same data as the lectotype and 1 ex. "Dare-dasht östlich Rudbar, 1000 m, 4.8.1968 ".

Additional material examined. TURKEY: Rize: 4 exs., ca. 30 km SW Hopa, Çaġlayan D. river valley, ca. 41°09'N, 41°22'E, 1800–1900 m, 26.VI.1998, leg. Solodovnikov (cSol, cAss); Artvin: 1 ex., ca. 40 km SW Artvin, source of Barhal river, ca. 41°05'N, 41°30'E, 2400 m, 23.VI.1998, leg. Solodovnikov (cAss); 2 exs., same data, but 2400–2800 m, 24.VI.1998 (cAss). GEORGIA: 1 ex., Caucasus minor, Trialetskiy Khrebet, Bakuriani, 1800–2200 m, 15.–20.VI.1987, leg. Schülke & Wrase (cSch); 3 exs., "Svanetien, Leder. Reitter" [labelled as paratypes of X. reitteri ] ( HNHM). AZERBAIJAN: 2 exs., Lenkoran, Apo below Bilasar, 350 m, 8.–9.VI.1996, leg. Schawaller ( SMNS, cAss); 1 ex., Gamarat [38°39'N, 48°35'E; "Caspi.-M.-Gebiet, Hamarat, Leder (Reitter)"], [labelled as paratype of X. reitteri ] ( HNHM); 1 ex., "Caspi.-M.-Gebiet, Rasano, Leder (Reitter)", [labelled as paratype of X. reitteri ] ( HNHM). IRAN: Mazandaran: 3 exs., Kalardasht, Rudbarak, 36°27'N, 51°04'E, 1790–1950 m, 26.VII.2005, leg. Frisch & Serri ( MNHUB, cAss); 3 exs., Kalardasht, Rudbarak, 36°28'N, 51°06'E, 1540–1640 m, 26.VII.2005, leg. Frisch & Serri ( MNHUB, cAss); 1 ex., Mazandaran, V.1999, leg. Reuter (cFel).

Comment. The original description of X. reitteri is based on a single holotype male from "Caucase, Swaneten [sic]" ( Coiffait 1966a, 1972). Consequently, the five specimens in the Reitter collection (HNHM) labelled as paratypes by Coiffait do not have type status. Xantholinus svanetskiensis was originally described based on a single holotype male from "Svanetskij Khrebet, Causace [sic]" ( Coiffait 1973).

There has been considerable confusion regarding the identities of X. reitteri , X. fortepunctatus Motschulsky, 1860 , and X. semirufus Reitter, 1901 (see discussion in section on X. fortepunctatus ). According to Bordoni (1975a), the interpretation of X. fortepunctatus by Coiffait (1972) and Korge (1973) is incorrect. An examination of the holotype of X. reitteri revealed that it is conspecific with X. fortepunctatus sensu Coiffait (1972) and Korge (1973). Bordoni (1999a), who designated a lectotype for X. fortepunctatus , referred Coiffait´s (1972) and Korge´s (1973) interpretation of X. fortepunctatus to a new species, X. (Helicophallus) motschulskyi , without designating a holotype and apparently also without examining the type material, but stating that the species was "nota al momento con sicurezza dell'Iran (Gilan)". Hence, the specimens from Gilan indicated by Korge (1973) have syntype status. One of the males from the type series is here designated as the lectotype of X. motschulskyi . Since the aedeagi of this specimen and that of the holotype of X. svanetskiensis are identical to that of the holotype of X. reitteri , it follows that both X. motschulskyi and X. svanetskiensis are junior synonyms of the senior name X. reitteri . According to the descriptions and illustrations in Coiffait (1966a, 1972), there are no spines in the distal part of the aedeagus. However, these can only be seen properly in lateral view and were probably overlooked. The aedeagus and its internal structures of X. reitteri are illustrated in Figs. 23–26 , that of the holotype in Fig. 98 . For comparison see also the illustrations of the aedeagus of X. fortepunctatus by Korge (1973), of X. reitteri and X. svanetskiensis by Coiffait (1972, 1973), and of X. reitteri by Bordoni (1975a).

Owing to the previous taxonomic confusion, most of the zoogeographic data in the literature pertaining to X. fortepunctatus are unclear and part of the records of X. fortepunctatus in fact refer to X. reitteri . Reliable records of X. reitteri were previously known from Georgia and Azerbaijan ( Bordoni 1975a, Coiffait 1966a, Ushakov 1988); according to Ushakov (1988), the species is widespread in the Caucasus region. It is here reported from Turkey (where it is probably confined to the extreme northeast) ( Fig. 22 ) and Iran for the first time.

Figures

FIGURE 22. Distributions of Xantholinus osellai Bordoni (open circles), X. grandespinosus Assing (filled circles), and X. reitteri Coiffait (squares) in Turkey.

FIGURES 23 – 30. Xantholinus reitteri Coiffait (23 – 26), X. sidonensis Coiffait, holotype (27), and X. rufipennis Erichson (28 – 30; 28 – 29: lectotype, 30: male from Cyprus): 23, 27, 29 – 30 — aedeagus in dorsal view; 24 — aedeagus in lateral view; 25 — distal internal structures of aedeagus; 26 — distal internal structures of aedeagus in lateral view; 28 — habitus. Scale bars: 28: 1.0 mm; 23 – 25, 27, 29 – 30: 0.5 mm; 26: 0.2 mm.

FIGURES 94 – 100. Xantholinus chersonesicus sp. n. (94 – 97), X. reitteri Coiffait, holotype (98), and Vulda ottomana (Cameron), holotype (99 – 100): 94, 99 — forebody; 95, 98, 100 — aedeagus; 96 — internal structures of aedeagus in squeeze preparation; 97 — distal part of internal structures of aedeagus in squeeze preparation. Scale bars: 94, 99: 1.0 mm; 95 – 98: 0.5 mm; 100: 0.2 mm.