Pterostichus makarovi, Fedorenko, 2023

Fedorenko, D. N., 2023, A review of Pterostichus subgenus Orientostichus (Coleoptera: Carabidae: Pterostichini) from Indochina, with description of new species, Russian Entomological Journal 32 (2), pp. 155-170 : 161

KEY TO SPECIES OF PTEROSTICHUS (ORIENTOSTICHUS)

FROM INDOCHINA.

1(10) Pronotal lateral bead fine, not or barely wider in basal than in apical half ( Figs 1–3 , 5–6 ). Pronotum narrow at base, its sides straight or subsinuate in front of basal angles; inner and outer basolateral sulcus joining along base. Elytron with seta d3, intervals mostly distinctly cross-striated on sides toward apex or densely transversely rugose in apical third. Metatibia with 1–4 setae along outer margin. Body large, BL 22–27 mm. ............... The prattii -group.

2(5) Elytra down the middle dull due to intervals densely transversely rugose ( Fig. 6 ); elytral microsculpture isodiametric. Pronotal basal angles pointed, right or acute.

3(4) Elytral intervals subequally developed. Pronotal base broader, PB/PA 1.29. — Northern Myanmar. ............... ................................. P. semirugosus ( Andrewes, 1947)

4(3) Odd elytral intervals more developed than even ones in apical half, with their median shiny stripes reaching apex or not, respectively. Pronotal base narrower, PB/PA 1.19. — Northern Thailand............. 4. P. ferreroi Straneo, 1989

5(2) Elytra shiny, with intervals more or less distinctly cross-striated toward apices ( Figs 10–11 ), or entirely dull from coarse isodiametric microsculpture in female. Pronotal basal angles more or less obtuse and mostly blunt.

6(7) Elytral microsculpture cisodiametric in both sexes, base of interval 8 in form of fairly long carina. Meso- and metatarsomeres 1–3 with dorsolateral setae. Abdominal sternite VII in male with a high median carina ( Fig. 14 ). — Southern China. ................... [3. P. prattii Bates, 1890 ]

7(6) Elytral microsculpture consisting of transverse meshes or transverse lines, or granulate in female. Meso- and metatarsomeres 1 and 2 with dorsolateral setae. Abdominal sternite VII in male with a small, slightly longitudinal, tubercle in and slightly down the centre ( Figs 15, 18 ). — Northern Vietnam.

8(9) Pronotal base densely punctate on each side. Elytral microsculpture sexually dimorphic, coarse and granulate (♀) or consisting of superficial transverse meshes (♂), thus giving elytra dull or shiny appearance, respectively. Reflexed lateral margin of elytra with a fine carina in apical half. Pronotal lateral bead wide, more so in basal half. — Phia Oac Mt. .................... 2. P. makarovi sp.n .

9(8) Pronotal base impunctate to finely and very sparsely punctate on each side. Elytra shiny, with microsculpture consisting of very transverse superficial meshes (♀) or dense transverse lines (♂). Reflexed lateral margin of elytra with no carina. Pronotal lateral bead narrow, barely wider in basal than in apical half. — Hoang Lien Son Mt. ridge. ........... ......................................................... 1. P. matalini sp.n .

10(1) Pronotal lateral bead thick anteriorly, flat and dilated in basal half ( Figs 4 , 7–9 ). Pronotum wide at base, rounded on sides; inner and outer basolateral sulci separate, basal bead missing. Elytron without discal setae, intervals smooth. Metatibia asetose at outer margin .................... .................................................... The lesticoides -group.

11(14) Body smaller, BL 18–20 mm. Elytra without discal setae, with microsculpture consisting of very dense transverse lines on disc and of longitudinal meshes at bottom of reflexed lateral margin. — Hoang Lien Son Mt. ridge.

12(13) Pronotal base wide, mean PB/PA~1.3. Aedeagus apex in lateral view with dorsal margin distinctly s-shaped ( Fig.4 ). ....................... 5. P. lesticoides ( Straneo, 1939)

13(12) Pronotal base narrow, mean PB/PA~1.1. Aedeagus apex in lateral view with dorsal margin smoothly sinuate ( Fig.7 ). ............................ 6. P. pseudolesticoides sp.n.

14(11) Body large, BL 22–23 mm. Elytra with discal seta d3 adjoining stria 2 a fourth from apex and microsculpture consisting of transverse meshes on disc and of isodiametric meshes at bottom of reflexed lateral margin. Pronotal base wide, PB/PA 1.27 or 1.30.

15(16) Elytral striae smooth, reflexed lateral margin with a slight and blunt carina in apical half. — Phia Oac Mt. ... .................................................. 7. P. laevistriatus sp.n.

16(15) Elytral striae finely yet distinctly punctate, reflexed lateral margin with a vague carina in apical half. — Tam Dao Son Zuong Mt. ridge. ....... 7. P. tamdaoensis sp.n.

Figures

Figs 1–3. Dorsal habitus: 1 — Pterostichus makarovi sp.n., paratype ♀; 2–3 — P. matalini sp.n., paratypes from Phan Xi Pang Mt or Bat Xat NP. Рис. 1–3. Габитус дорЗально: 1 — Pterostichus makarovi sp.n., паратип ♀; 2–3 — P. matalini sp.n., паратипы с г. Фансипан и иЗ Национального парка Батсат.

Figs 4–6. Dorsal habitus: 4 — Pterostichus lesticoides, holotype; 5 — P. prattii, ♀ from Chengkou; 6 — P. ferreroi, paratype ♂. Рис. 4–6. Габитус дорЗально: 4 — Pterostichus lesticoides, голотип; 5 — P. prattii, ♀ иЗ Chengkou; 6 — P. ferreroi, паратип ♂.

Figs 10–18. Pterostichus: 10, 15 — P. matalini sp.n.; 11, 14 — P. prattii, from Moupin, China; 12 — Pterostichus sp. from China; 13 — P. curtatus; 16 — P. pseudolesticoides sp.n.; 17 — P. ferreroi; 18 — P. makarovi sp.n.; 10–12 — apices of elytra and abdomen, dorsal aspect; 13–18 — abdominal sternite VII in male, left lateral view. Not to scale. Рис. 10–18. Pterostichus: 10, 15 — P. matalini sp.n.; 11, 14 — P. prattii, иЗ Moupin, Китай; 12 — Pterostichus sp. иЗ КитаЯ; 13 — P. curtatus; 16 — P. pseudolesticoides sp.n.; 17 — P. ferreroi; 18 — P. makarovi sp.n.; 10–12 — вершины надкрылий и брюшка, дорЗально; 13–18 — стернит VII брюшка саца, вид слева. БеЗ масштаба.

Figs 7–9.Dorsal habitus:7 — P. pseudolesticoides sp.n., paratype ♀; 8 — P. laevistriatus sp.n., holotype; 9 — P. tamdaoensis sp.n., holotype. Рис. 7–9. Габитус дорЗально: 7 — P. pseudolesticoides sp.n., paratype ♀; 8 — P. laevistriatus sp.n., голотип; 9 — P. tamdaoensis sp.n., голотип.