Scaphisoma biplagiatum HELLER, 1917

Ivan Löbl & Ryo Ogawa, 2016, On the Scaphisomatini (Coleoptera, Staphylinidae, Scaphidiinae) of the Philippines, IV: the genera Sapitia ACHARD and Scaphisoma LEACH, Linzer biologische Beiträge 48 (2), pp. 1339-1492 : 1399-1400

5.2.14.3. Scaphisoma biplagiatum HELLER, 1917 (Figs 8, 144, 145)

Scaphosoma biplagiatum HELLER, 1917 : 47. Lectotype 6, SMTD; type locality: Luzon, Mt. Makiling. Additional published localities: Masbate: Aroroy, Luzon: Imugan, Bucas: Sorocco ( LÖBL 1972: 107).

Pseudoscaphosoma implagiatum PIC, 1926 : 1. Lectotype ♀, MNHN; type locality: Luzon, Bagnio. Characters: LÖBL 1970a: 126, 127, Fig. 2 . Synonymy and lectotype designation: LÖBL 1972: 107. Additional material examined: Luzon, Lagunas Prov., Mt. Makiling, summit rd., 600 m, 21-22 and 28.XI.1995, leg. I. Löbl, 103 ex. (MHNG); Luzon, Mt. Makiling, above Mad Springs, 400-700 m, degraded rainforest, 19-22.XI.1995, leg. J. Kodada, 10 ex. (MHNG); Luzon, Mt. Makiling, 4 km SE Los Banos, 9.IV.1977, leg. L.E. Watrous, 3 ex. (MHNG); Mt. Makiling, 17-18.VI.1977, leg. M. Satô, 1 ex. (MHNG); Luzon, Laguna Prov., Mt. Banahaw, above Kinabuhayan, 600-700 m, degrad. rainforest, 24.XI.1995, leg. I. Löbl, 2 ex. (MHNG); Luzon, Benguet Prov. Pellwell Cr., camp two, 14.IV.1977, leg. M. Satô, 1 ex. (MHNG); Luzon, Ifugao Prov., Mt. Polis, 1900 m, 4-5.VI.1977, leg. M. Satô, 1 ex. (MHNG); Palawan, Central, Conception, sea level, 6-7.XII.1995, leg. I. Löbl, 52 ex. (MHNG, BZLA); Palawan, Conception, large logs across Conception River, NE San Rafael,ca 20 m 8.XII.1995, leg. J. Kodada & B. Rigová, 25 ex. (MHNG); Palawan, Olanguan, 18 km NE San Rafael, sea level, 5-6.XII.1995, leg. I. Löbl, 106 ex. (MHNG); Palawan, Olanguan River, 18 km NE San Rafael, 6.XII.1995, leg. J. Kodada & B. Rigová, 1 ex. (MHNG); Palawan, Olanguan, 0-50 m, between Puerto Princesa and Roxas, 1.IX.1985, leg. M. Sakai, 9 ex. (EUMJ); Palawan, above San Rafael, ca 300 m, degraded forest on slope, 4.XII.1995, leg. J. Kodada, 1 ex. (MHNG); along Tarabanan River, NE San Rafael, ca 30 m, 7.XII.1995, leg. J. Kodada & B. Rigová, 2 ex. (MHNG); Palawan, Sabang, trail to Underground River, sea level, 30.XI.1995, 2 ex. (MHNG); Sabang, N. of Mt. St. Paul, 11- 13.VI.1977, leg. M. Satô, 2 ex. (MHNG); Palawan, Matalangao nr. Roxas, 150 m, 29.VIII.1985, leg. M. Sakai, 2 ex. (EUMJ); Palawan, Taytay Prov., Binaluan, 1 ex. (SMTD); Palawan, Narra Prov., Trident Mine, 100 m, Peak nr. Narra, 5.IX.1985, leg. M. Sakai, 1 ex. (EUMJ); Leyte, Visca N Baybay, sec. forest, 100-200 m, 27.II.1991, leg. W. Schawaller et al., 2 ex. (SMNS, MHNG); Mindanao, Baracatan, 1500 m, 27-29.1077, leg. M. Satô, 11 ex. (MHNG); Mindanao, 15 km S of Bislig, 3.VII.1977, leg. M. Satô, 1 ex. (MHNG); Mindanao, Todaya, 29.VII.1970, leg. M. Satô, 1 ex. (MHNG); Mindanao, Maitum, 150 m, South Cotabato Prov., 10.VIII.1985, leg. M. Sakai, 4 ex. (EUMJ); Mindanao, Matum Hot Springs, 28.VII.1970, leg. M. Satô, 1 ex (MHNG); Mindanao, Cotabato Prov., 50 km N. of Parang, 500 ft., 6.XII.1946, on encrusting polypore, 2 ex., leg. F.G. Werner (FMNH); Mindanao, Cotabato Prov., Burungkot, Upi, 1500 ft., 1-9.I.1947, on polypores, 18 ex., leg. F.G. Werner (FMNH, MHNG); Mindanao, Davao Prov., E. slope of Mt. McKindley, 3200 ft., 7-8.IX.1946, beating, 2 ex., leg. F.G. Werner (MHNG); Mindanao, 30 km E of Malaybalay, Busdi, 1000 m, 5-9.V.1996, leg. Bolm, 6 ex. (SMNS); Mindanao, Misamis occ., Don Victoriano, 1700 m, 1-3.V.1996, leg. Bolm, 5 ex. (SMNS); Mindanao, Mt. Apo, Ilomavis, 1400 m, 18-19.V.1996, leg. Bolm, 1 ex. (SMNS); Mindanao, 25 km W Zamboanga, 800 m, Camp Susana, 29-30.V.1996, leg. Bolm, 1 ex. (SMNS).

Distribution: Philippines: Luzon, Palawan, Masbate, Bucas Grande, Leyte, Mindanao.

Comments: The aedeagal characters illustrated in LÖBL 1970a: 127 were taken from the sole specimen available at that time, a male with partly extruded internal sac, and lack in detail. Therefore, new illustrations are given in the present work ( Figs 144, 145 ). The species is a member of the S. pictum group, characterized by the internal sac with a large basal tuft of spines joined with a large basal tooth, in combination with a large apical denticle and a subrectangular or subpentagonal central sclerite. The aedeagal characters suggest close relationship with S. dohertyi PIC, 1915, widely distributed in tropical Asia. The basal bulb of the median lobe is extended in S. dohertyi, unlike that in S. biplagiatum, and completely overlapping the apical process of the median lobe.

Figures

Figs 1 - 6: Habitus of Scaphisoma; S. anomalum LÖBL (1); S. bibiranense LÖBL (2); S. bicolor nov. sp. (3); S. bicoloripenne nov. sp. (4); S. bicuspidatum nov. sp. (5); S. binaluanum PIC (6). Scale: 1.0 mm.

Figs 142 - 145: Aedeagi of Scaphisoma in dorsal and lateral views; (142, 143), S. bicoloripenne nov. sp., scale = 0.2 mm; (144, 145) S. biplagiatum HELLER, scale = 0.1 mm.