Nothomydas namaquensis Hesse

Dikow, T., 2012, Review of Namibimydas Hesse, 1972 and Nothomydas Hesse, 1969 (Diptera: Mydidae: Syllegomydinae: Halterorchini) with the description of new species, African Invertebrates 53, pp. 79-111 : 100-101

Nothomydas namaquensis Hesse ,1972

Figs22, 29-31, 39, 40, 43

Nothomydas namaquensis :Hesse1972:152.

ZooBank LSID:see Table1.

Diagnosis:The species is distinguished from congeners by the predominantly grey pubescent scutum with apubescent anterior slopes,the entirely grey pubescent scutellum,the dorsally and ventrally white setose scape and pedicel,the structure of the postpedicel,which is cylindrical in the proximal quarter and bulbous in the distal three quarters,and its apparent distribution in coastal habitats in the north-western Northern Cape Province of South Africa(Fig.43).

Redescription:

Male.

Head:Brown,in general densely grey pubescent;width distinctly greater than thorax,interocular distance on vertex larger than at ventral eye margin,vertex between compound eyes more-or-less horizontally straight,medially only slightly below dorsal eye margin,parafacial area about as wide as half the width of central facial gibbosity;facial gibbosity distinct,well-developed and discernible in lateral view;mystax white,covering entire facial gibbosity;frons medially apubescent,laterally grey pubescent,vertex medially apubescent,laterally grey pubescent,postgena apubescent;setation:vertex white,frons white,ocp setae white,pocl macrosetae absent;ocellar triangle apubescent;proboscis brown,long,projecting well-beyond fronto-clypeal suture and beyond tip of postpedicel;labellum small,as wide as prementum,about quarter length of prementum,unsclerotised laterally;maxillary palpus cylindrical,brown,minute.

Antenna:Brown,scape and pedicel white setose dorsally and ventrally;postpedicel cylindrical in proximal quarter,symmetrically bulbous in distal three-quarters,≥6.0 × as long as combined length of scape and pedicel;apical ‘seta-like’ sensory element situated apically in cavity on postpedicel.

Thorax:Brown,scutum predominantly grey pubescent,pleura predominantly apubescent;scutum uniformly brown,surface entirely smooth,anterior slopes(facing occiput)apubescent,otherwise grey pubescent with broad sublateral stripes(not reaching anterior margin)and narrower paramedian stripes(not reaching posterior margin)brown pubescent,scutal setation comprised of long white setae with distinct rows of long dorsocentral setae and dense lateral scutal setae;dc setae pre- and postsuturally white,acr setae present,lateral scutal setae white,npl setae0,spal setae0,pal setae0;postpronotal lobe light brown,grey pubescent;proepisternum,lateral postpronotum,and postpronotal lobe long white setose;scutellum grey pubescent,asetose,apical scutellar setae absent;mesopostnotum,anatergite,and katatergite grey pubescent,mesopostnotum asetose,anatergite asetose,katatergite long white setose;katatergite more-or-less flat;anterior anepisternum asetose,supero-posterior anepisternum long white setose;posterior anepimeron long white setose,katepimeron long white setose;metanepisternum grey pubescent,asetose,metepimeron more-or-less flat,same colour as T1,grey pubescent,long white setose.

Leg:Brown,setation white on pro and mes legs and white and brown on met leg;pro,mes,and met coxa apubescent,pro and mes coxa short white setose,met coxa long white setose;met trochanter setose medially;femur brown,met femur more-or-less cylindrical only slightly wider than pro and mes femur,in distal half macrosetose,1anteroventral and1posteroventral row of macrosetae,posteroventrally long white,erect setose and setae arranged in distinct row;pro,mes,and met tibia straight,met tibia cylindrical,ventral keel absent,lateroposteriorly long white,erect setose and setae arranged in distinct row;pro and mes tarsomere1longer than tarsomere2,but less than combined length of tarsomeres2-3,met tarsomere1less than combined length of tarsomeres2-3;pulvillus well-developed on pro and mes legs,smaller on met legs;empodium absent.

Wing:Length 5.5-6.1mm;hyaline throughout,veins brown,microtrichia absent;cells r1,r4,r5,m3,and cup closed(r5sometimes open);C terminates at junction with M1(or M1+M2)or terminates at junction with R1;R4terminates in R1;R5terminates in R1;stump vein(R3)at base of R4absent;R4and R5widest apart medially; r–m distinct,R4+5and M1apart,connected by crossvein;M1straight at r–m (not curving anteriorly),M1(or M1+M2)terminates in C or R1;CuA1and CuA2split proximally to m–cu (cell m3narrow proximally);M3+CuA1do not terminate together in C;A1undulating,cell a1wide,A1and wing margin further apart proximally than distally;alula well-developed;haltere light brown.

Abdomen:Brown;setation comprised of dense white and brown setose,surface entirely smooth;T1-7brown;T1and anterior half of T2long white setose,remaining T2and T3-7brown setose;T1apubescent,T1-7anterior0.25and laterally apubescent,remaining T grey pubescent;S1-7brown;S1asetose,S2-3white and brown setose;S mainly apubescent;T2-4parallel-sided and not constricted waist-like;bullae on T2black,transversely elongate,surface entirely smooth,T2surface anterior to bullae smooth.

Terminalia(Figs29-31):Epandrium distally in dorsal view pointed with bid tip posteromedially.

Female.

Head:Mystax white,covering facial gibbosity except for narrow median dorso-ventral area.Antenna:Brown;scape and pedicel white setose dorsally and ventrally;postpedicel cylindrical in proximal quarter,symmetrically bulbous in distal three-quarters,≥6.0 × as long as combined length of scape and pedicel;apical ‘seta-like’ sensory element situated apically in cavity on postpedicel.Leg:Setation predominantly white;in distal half macrosetose,1anteroventral row of macrosetae.Wing:Length 6.1mm;veins light brown;C terminates at junction with R;M(or M+M)terminates in R.1 1 1 2 1Abdomen:Scattered white setose,T1and anterior quarter of T2long white setose,remaining tergites short white setose,setae directed anteriorly on T3-7;S1asetose,S2-7sparsely white setose.Genitalia(Fig.22):5-6acanthophorite spines per plate.

Material examined: SOUTH AFRICA: Northern Cape: 1♀ 5♂ Port Nolloth , 29°15'17"S 16°52'18"E, x.1964, SAM Museum Staff (holotype & paratypes, SAMC) .

Type locality and distribution:Port Nolloth(29°15'17"S 16°52'18"E),South Africa(Fig.43,GBIF resource#14003).Biodiversity hotspot:endemic to Succulent Karoo.