Manota abscissa, Hippa & Ševčík, 2013

Hippa, Heikki & Ševčík, Jan, 2013, Five new species and a new record of Manota (Diptera: Mycetophilidae) from Sulawesi, Acta Entomologica Musei Nationalis Pragae 53 (2), pp. 763-775 : 764-766

Manota abscissa sp. nov.

( Figs 1A–D )

Type material. HOLOTYPE: J, INDONESIA: SOUTH SULAWESI: ‘ Bulusaraung forêt secondaire altitude, au sol, Site 2-1, Grand Malaise 3, 18– 27.8.2007, INDO710 M1B’ ( MNHN) . PARATYPES: 3 JJ with the same data as holotype ; 1 J, INDONESIA: SOUTH SULAWESI: ‘ Bulusaraung, forêt secondaire altitude, 04º55.861′S, 119º45.369′E, 1066 m, Site 1-1, Grand Malaise 1, 18– 27.8.2007, INDO708 M1B’ ( MNHN) .

Description of male. Colour. Head brown, face and clypeus paler brown. Antenna brown or scapus and pedicellus paler brown. Mouth parts yellowish. Thorax pale brown, preepisternum 2 ventrally paler. Legs yellowish, coxa 3 infuscated at base. Wing pale brownish. Halter pale brown with blackish knob. Abdomen brown, sternites paler than tergites. All setosity pale, yellowish or brownish. Head. Antennal flagellomere 4, Fig. 1A . Palpomere 3 of maxillary palpus with apicomesial thumb-like extension, with 4–5 apically curved sensilla; palpomere 4 with parasegment; palpomere 5 ca. 1.3 times longer than palpomere 4. Number of strong postocular setae 10–11. Thorax. Anepisternum setose, with 42–54 setae; anterior basalare non-setose; preepisternum 2 setose, with 15–18 setae; laterotergite setose, with 18–27 setae; episternum 3 setose, with ca. 10 setae. Legs. Mid and hind tibial organs absent. Wing. R1 meeting C well on the basal half of the costal margin; the sclerotized part of M2 extending near to the level of the tip of R1; wing length 1.6–1.8 mm. Hypopygium ( Figs 1B–D ). Lateral margin of tergite 9 free from the gonocoxa, convex, posterior part submembranous, transverse, not quite reaching the middle of gonocoxa, anterior margin deeply incised, the setae similar to the ventral setae of gonocoxa. Ventral mesial margin of gonocoxa simple; the ventral setae of gonocoxa unmodified, those at the mesial margin long. Parastylar lobe subtriangular, with two setae at the anterior narrower part. Paraapodemal lobe large. Dorsal mesial margin of gonocoxa simple, slightly bulging towards the medial line on the posterior half, posteriorly with a weak setose lobe. Gonocoxa without any posterolateral lobe. The dorsal setae of gonocoxa unmodified, similar to those on the ventral side. Two juxtagonostylar megasetae, both are similar long, strong megasetae, the ventral one apically curved, the dorsal one apically almost straight, both arising from a common basal body which is ca. one fourth of the length of the megasetae. No other setae on the dorsal side of gonocoxa that deviate from the general setosity. Gonostylus large, elongate. Ventral side of gonostylus with very long setae, except mesially on the basal half non-setose, dorsal side almost non-setose except for a few short setae subbasally and subapically. Aedeagus elongate subtriangular, with weak lateral shoulders. Hypoproct extending posteriorly to the base of gonostyli, number of its ventral setae (sternite 10) ca. 6 on each half, arranged in a row. Cerci mesially separate.

Differential diagnosis. Setose anepisternum, setose preepisternum 2, setose laterotergite, simple subtriangular, not apically deeply bilobed or unusually elongated aedeagus and medially separate cerci combine Manota abscissa sp. nov. with five other Oriental species: M. aciculata sp. nov., M. angustata Hippa, 2006 , M. biunculata Hippa, 2007 , M. inusitata Hippa & Papp, 2007 and M. ovata Hippa, 2006 . It is distinguished from M. aciculata sp. nov. and M. inusitata by lacking megasetae in any position on its gonostylus. It is distinguished from M. ovata by having the ventral setae on the hypoproct arranged in a row on both halves, not in a patch, and by having the mesial marginal area on the basal half of the gonostylus non-setose (with long setae in M. ovata ). It is distinguished from M. angustata by lacking a prominent setose lobe apicomesially on the gonostylus.

Etymology. The name is a Latin adjective, abscissus, -a, -um, meaning cut off, referring to the dorsal one of the juxtagonostylar megasetae, which seems as if being cut off at the apex.

Figures

Fig. 1. Manota abscissa sp. nov. (holotype).A – antennal flagellomere 4, lateral view. B – hypopygium, dorsal view. C – hypopygium, ventral view. D – hypoproct and aedeagus, ventral view. Scales 0.10 mm. 1 = sternite 9, 2 = gonocoxa, 3 = ventral mesial margin of gonocoxa, 4 = parastylar lobe, 5 = paraapodemal lobe, 6 = gonostylus, 7 = apex of aedeagus, 8 = aedeagal apodeme, 9 = juxtagonostylar setae, 10 = dorsal mesial margin of gonocoxa, 11 = lobes at dorsal posterior margin of gonocoxa, 12 = gonocoxal apodemes, 13 = tergite 9, 14 = hypoproct, 15 = cercus.