Glypturus toulai ( Rathbun, 1919 )

Klompmaker, A., Hy ̆ z, Matu ̆ s, , Roger W. Port & Kowalew, nd Michał, 2016, Growth, inter- and intraspecific variation, palaeobiogeography, taphonomy and systematics of the Cenozoic ghost shrimp Glypturus, Journal of Systematic Palaeontology 14 (2), pp. 99-126 : 116

Glypturus toulai ( Rathbun, 1919)

( Fig. 16)

‡ 1919 Callianassa toulai Rathbun , 146.

2013 Glypturus toulai (Rathbun) ; Hy̆zńy et al.: 132, figs 4, 5 (cum syn.).

Diagnosis. Moderately tuberculated Glypturus ; lateral tuberculation on outer side of propodus extending from near base of fixed finger diagonally to lower margin and reaching proximal lower corner; lateral tuberculation on inner side of the propodus restricted mainly to area of articulation with dactylus in small specimens, but covering [central portion of inner side of manus in large specimens.] (modified from Hy̆zńy et al. 2013, pp. 132 — 133).

Material. UF 203348 ( Fig. 16E, F). NHMW 1933/0018/ 0159 ( Fig. 16A — D), 1933/0018/0160 ( Fig. 16G — J + 3 fixed fingers). BMNPH PI IC 395 ( Fig. 16K — M).

Occurrence. This species is known from the upper Miocene Gatun Formation in Panama (Supplemental Table 1 ).

Description. Manus length exceeding height (length/ height ratio ~1.1). Upper margin converging distally, proximally keeled, bearing three prominent spines pointing distally, proximalmost spine around mid-margin, keel terminating in blunt corner. Lower margin sharp and keeled, in larger specimens denticulate, lined with a row of setal pits on inner lateral side. Proximal margin convex on outer face, concave on inner, [heightwise groove near proximal margin concave on inner side, concave to straight on outer side]. Distal margin weakly convex. Lateral tuberculation on outer side from base of fixed finger diagonally to lower margin and proximal lower corner. Lateral tuberculation on inner side of propodus restricted mainly to area of articulation with dactylus in small specimens, but covering [central portion of inner side of manus in large specimens]. Fixed finger triangular, sharply pointed, with distinct blunt tooth on occlusal margin around mid-length, tooth pointing distally. Dactylus stout, occlusal margin [with or] without tooth [around mid-length], tip hooked (modified from Hy̆zńy et al. 2013, pp. 134 — 135).