Alaolacon megalopus, Arimoto, Koichi & Arimoto, Hisayuki, 2017

Arimoto, Koichi & Arimoto, Hisayuki, 2017, The genus Alaolacon Candeze, a senior synonym of the genus Eumoeus Candeze (Coleoptera, Elateridae, Agrypninae), ZooKeys 656, pp. 85-110 : 95-98

Alaolacon megalopus sp. n. Figures 38-40, 41-48, 49-54

Eumoeus murrayi Candèze, 1874; Fleutiaux 1928: 178 (mention a specimen from Java at IRSNB); Casari-Chen 1993: 241 (examined a male specimen from Java). Misidentification.

Etymology.

A combination of the Greek megalos, meaning great, and the Greek ops, meaning eye, refer to the large compound eyes.

Type material.

Holotype. Male, Java, Indonesia [IRSNB] (Fig. 40).

Diagnosis.

Body brown, without metallic luster; setae yellow-brown; frontal depression moderate; eye very large; prothorax wider than long, widest posteriorly; pronotal anterior angles rounded; median longitudinal depression shallow, located at anterior half, punctate; anterior angles of hypomeron acute; prosternal spine inclined strongly behind procoxae; scutellum 1.5 × as long as wide; sides of scutellum parallel; hind wings with cross vein between veins MP4 and CuA2 located just anterior to contact point between veins MP3 and MP4, without wedge cell; male tergite VIII longer than wide; median lobe of male aedeagus elongate.

Measurements.

BL: 11.8, BW: 3.54, MIE: 1.02, MAE: 2.31, OI: 44, PL: 3.11, PML: 2.74, PW: 3.26, PI: 95, EL: 7.70, EW: 3.54, EI: 218.

Description.

Body (Figs 38, 39) shining, without metallic luster. Color. Body brown; antennomere I, pronotal lateral margin, elytra, legs, abdomen paler; antennomeres II-XII, mouth-parts, pregenital segments and aedeagus yellow-brown, but mandible brown. Hairs. Body covered with yellow-brown setae; antennomeres III-XII with short filiform setae at ventral surface.

Head (Fig. 41). Frontal depressed moderate. Eyes very large. Antennomere I elongate; antennomere II short and obconical; antennomeres III-X flabellation moderate (Fig. 42). Apical maxillary palpomere (Fig. 43) rounded, 1.3 × as long as wide. (Antennomeres XI-XII of right antenna and antennomeres III-XII of left antenna lost.)

Prothorax wider than long; sides widest posteriorly, rounded anteriorly, liner posteriorly. Pronotum convex; anterior angles rounded; median longitudinal depression shallow, located at anterior half, punctate (Fig. 41); central area with two shallow concaves. Prosternal spine with lateral margin of dorsal side expanded laterally, inclined strongly (at 15 degrees) behind procoxae (Fig. 44). Hypomeron with anterior angles acute (Fig. 45); punctures more homogeneous than prosternal punctures in density and size. Scutellum (Fig. 47) 1.5 × as long as than wide; sides parallel. Elytra with sides almost parallel on basal half; intervals with small and coarse punctures. Hind wings (Fig. 48) with veins posterior to MP3 translucent; cross vein between veins MP4 and CuA2 not completely connected with CuA2, located just anterior to contact point between veins MP3 and MP4; wedge cell absent (Fore legs except for coxae, tarsomeres IV-V and claw of right middle leg, tarsomere V and claw of left middle leg, tarsi and claw of right hind leg, and left hind leg lost.)

Abdomen. Ventrite V 0.65 × as long as wide. Tergite VIII (Fig. 49) 1.2 × as long as wide; basal area translucent. Sternite VIII (Fig. 50) with central area translucent; median notch shallow and truncate transversally. Tergite IX (Fig. 51) with median notch shallow and rounded. Tergite × (Fig. 51) posterior margin rounded. Sternite IX (Fig. 52) wide; posterior half abruptly narrowed to apex; posterior margin rounded. Aedeagus (Figs 53, 54). Median lobe elongate, basal struts 0.37 × total length of median lobe. Parameres with sparse and short setae. Basal piece 0.28 × total length of aedeagus.

Female.

Unknown.

Distribution.

Indonesia: Java.

Remarks.

The holotype is damaged with most appendages lost. The holotype of this species was identified as Eumoeus murrayi (= Alaolacon murrayi comb. n.) by Candèze (Fleutiaux, 1928), but separated from Alaolacon murrayi by the following characteristics (the holotype of Alaolacon murrayi in parentheses): eye very large (OI: 44) (eye large, OI: 50); anterior angles of pronotum rounded (Fig. 38) (anterior angles of pronotum bisinuate, Fig. 56); pronotal median longitudinal depression shallow, located at pronotal anterior half and punctate (Fig. 41) (pronotal median longitudinal depression not reaching anterior margin or base and impunctate at posterior half); scutellum 1.5 × as long as wide (Fig. 47) (scutellum 1.3 × as long as wide, Fig. 56); scutellum sides parallel (Fig. 47) (scutellum sides concave and widest posteriorly, Fig. 56); hind wings with cross vein between veins MP4 and CuA2 (Fig. 48) (hind wings without cross vein between veins MP4 and CuA2); male tergite VIII longer than wide (Fig. 49) (male tergite VIII shorter than wide).

Only this species exhibits parallel sides of scutellum in this genus. The scutellum shape could be a useful specific diagnostic feature for this species including its female.