Stenus ater Mannerheim, 1830

Serri, Sayeh & Frisch, Johannes, 2016, Species diversity, chorology, and biogeography of the Steninae MacLeay, 1825 of Iran, with comparative notes on Scopaeus Erichson, 1839 (Coleoptera, Staphylinidae), Deutsche Entomologische Zeitschrift 63 (1), pp. 17-44 : 19

Stenus ater Mannerheim, 1830 Fig. 3 , Suppl. material 1

Chorology.

Stenus ater is widespread in the West Palaearctic and recorded as far east as European Russia, Azerbaijan, and North Iran, where it reaches its eastern limit of distribution in the Turkmeno-Khorasanian Mountains at about 58°06'E (Fig. 3 ). Records from Kazakhstan ( Smetana 2004: 554), which were adopted by Schülke and Smetana (2015: 806), mainly refer to other species and are not reliable ( Kastcheev and Puthz 2011: 454). In Iran, Stenus ater is the most widespread member of the Stenus ater group. Towards the east, it is replaced by its close relative Stenus mongolicus . The two species occur sympatrically in the eastern Elburz and the Turkmeno-Khorasanian Mountains (Fig. 3 ). Ghahari et al. (2009b: 1964) recently published Stenus ater for Behshahr, Mazandaran, a record which is in accord with the distribution pattern of the species.

Biogeographical characterization.

Judging from the vast distribution in the Mediterranean, we consider Stenus ater to be an expansive Holomediterranean faunal element.

Figures

Figures 1 - 6. Distribution of Steninae in Iran. 1. Geographical subdivision of Iran modified after Petrov (1955) and 2 - 6 distribution data of Steninae: 2. Dianous and the Stenus alpicola species group: ● Dianous coerulescens korgei, ○ Stenus proprius; 3. Stenus ater species group: ➕ Stenus affaber, ● Stenus ater, △ Stenus hypoproditor, ■ Stenus intricatus zoufali, ● Stenus mongolicus, ○ Stenus nodipes, ▲ Stenus skoraszewskyi; 4. Stenus brunnipes species group: ● Stenus brunnipes brunnipes; 5. Stenus canaliculatus species group: ● Stenus canaliculatus; 6. Stenus cautus species group: ○ Stenus cautus. Abbreviations: C - Caspian wet forest region, D - deserts and salt marshes, G - Golestan region, Gh - Ghohrud Mountains, Hm - mountains of Hormozgan, Khu - Khuzestan Plain, NB - North Baluchestan Mountains, NKh - North Khorasan Mountains, NNW - northern part of North Zagros Mountains, NZ - North Zagros Mountains, SB - South Baluchestan Mountains, SC - coast of Persian Gulf and Oman Sea, SE - southern slopes of Elburz Mountains, SKh - southern Khorasan Mountains, SZ - southern Zagros Mountains in Fars, Z - Zagros Mountains.