Jocquestus, Lyle & Haddad, 2018

Lyle, Robin & Haddad, Charles R., 2018, Jocquestus, a new genus of trachelid sac spiders from the Afrotropical Region (Arachnida: Araneae), Zootaxa 4471 (2), pp. 309-333 : 310-312

publication ID

https://doi.org/ 10.11646/zootaxa.4471.2.4

publication LSID

lsid:zoobank.org:pub:691AD8B9-27BB-40E6-A3D8-C3D17DA38B0B

DOI

https://doi.org/10.5281/zenodo.5949501

persistent identifier

https://treatment.plazi.org/id/F3ECF685-9890-4E9E-947B-DE5B952357DB

taxon LSID

lsid:zoobank.org:act:F3ECF685-9890-4E9E-947B-DE5B952357DB

treatment provided by

Plazi

scientific name

Jocquestus
status

gen. nov.

Genus Jocquestus gen. nov.

urn:lsid:zoobank.org:act:F3ECF685-9890-4E9E-947B-DE5B952357DB

Type species. Trachelas schenkeli Lessert, 1923

Etymology. The genus name is a patronym in honour of Rudy Jocqué, former head of the Invertebrate Section of the Royal Museum for Central Africa in Tervuren, Belgium, in recognition of his vast contributions to the

systematics of spiders, particularly the Zodariidae , Corinnidae and Ctenidae of the Afrotropical Region, and to arachnology in general. Gender masculine.

Diagnosis. Jocquestus gen. nov. can be recognized from other genera of smaller trachelid spiders, particularly Trachelas and the Afrotropical Fuchiba , Fuchibotulus and Planochelas , by males having blunt ventral cusps on the tibiae and metatarsi of leg I, and sometimes also on tarsi I and/or tibiae and metatarsi II ( Figs 7 View FIGURES 1–8 , 30–32 View FIGURES 21–32 ), which are absent in females ( Fig. 8 View FIGURES 1–8 ). Males of Jocquestus gen. nov. share with the Eurasian genus Paratrachelas the ventral cusps at least on leg I of males, but can be distinguished by the presence of a large RPA, and absence or presence of a short blunt RTA on the male palpal tibia (e.g. Figs 37 View FIGURES 37–40 , 49 View FIGURES 47–51 ). In Paratrachelas the RPA is entirely absent, while the RTA is strongly developed (see Kovblyuk & Nadolny 2009). Females are very distinct amongst these aforementioned genera of smaller trachelines by the heavily sclerotized epigastric plate, with broad, generally obliquely-directed epigynal ridges containing lateral copulatory openings ( Fig. 40 View FIGURES 37–40 ).

Description. Small spiders, 2.20–4.10 mm in length. Carapace: orange-brown, red-brown or dark brown ( Figs 1, 2 View FIGURES 1–8 , 9–20 View FIGURES 9–20 ); broadly oval with narrow eye region, widest at midpoint ( Figs 1, 2 View FIGURES 1–8 , 21 View FIGURES 21–32 ); dome-shaped in lateral profile, elevated from clypeus to behind posterior eyes, highest at midpoint, with steeper slope in posterior third ( Figs 3, 4 View FIGURES 1–8 , 22 View FIGURES 21–32 ); surface finely wrinkled, sparsely covered in short straight setae with finely granulate bases ( Figs 21–24 View FIGURES 21–32 ); posterior margin nearly straight, very slightly concave ( Fig. 21 View FIGURES 21–32 ); clypeus height approximately equal to AME diameter ( Fig. 23 View FIGURES 21–32 ). Eyes: AER slightly recurved in dorsal view ( Fig. 21 View FIGURES 21–32 ), procurved in anterior view, AME smaller than ALE ( Fig. 23 View FIGURES 21–32 ); AME closer to ALE than to each other; PER recurved in dorsal view, PLE very slightly larger than PME ( Fig. 24 View FIGURES 21–32 ), or equal in diameter; PME closer to each other than to PLE; all eyes surrounded by black rings; chilum single, retracted, triangular, rounded or with slight notch distally; cheliceral promargin and retromargin each with one to three teeth; cheliceral escort seta (sensu Ramírez 2014) present in front of fang base ( Fig. 25 View FIGURES 21–32 ); labium trapezoidal; endites depressed laterally, with distinct serrula ( Figs 26, 27 View FIGURES 21–32 ). Sternum and associated sclerites: sternum shield-shaped, strongly rebordered ( Figs 5, 6 View FIGURES 1–8 ); pleural bars isolated from each other and intercoxal sclerites; precoxal triangles present; intercoxal sclerites present between all coxal pairs. Legs: legs I, II and IV nearly equal in length, leg formula usually 1243, rarely 2143, 4123 or 4132; anterior legs more strongly built than posteriors, particularly in males ( Figs 1, 2 View FIGURES 1–8 , 9–20 View FIGURES 9–20 ); legs without spines, all segments covered with short fine setae, with scattered long erect setae on femora and tibiae I ( Figs 7, 8 View FIGURES 1–8 ); femora I swollen, strongly constricted proximally, more distinctly in males ( Fig. 7 View FIGURES 1–8 ) than females ( Fig. 8 View FIGURES 1–8 ); patellar indentation narrow, with broad lyriform organ at proximal end ( Figs 28, 29 View FIGURES 21–32 ); anterior tibiae, metatarsi and tarsi with ventral cusps in males ( Figs 7 View FIGURES 1–8 , 30–32 View FIGURES 21–32 ), absent in females ( Fig. 8 View FIGURES 1–8 ), with pseudotenent setae ventrally on tarsi only ( Fig. 33 View FIGURES 33–36 ); metatarsi III and IV with dense terminal preening brush; tarsal claws short, paired, claw tuft plates with dense claw tufts comprising tenant setae ( Fig. 33 View FIGURES 33–36 ); tarsi with sparse chemosensory setae ( Fig. 34 View FIGURES 33–36 ) and trichobothria ( Fig. 35 View FIGURES 33–36 ) dorsally; tarsal organ only very slightly elevated from surface, with faint ridges and oval opening ( Fig. 36 View FIGURES 33–36 ). Abdomen: broadly oval, tapering posteriorly, grey dorsally, occasionally with cream markings ( Figs 1, 2 View FIGURES 1–8 , 9–20 View FIGURES 9–20 ); male with dorsal scutum covering entire abdomen ( Fig. 1 View FIGURES 1–8 ), only present in females of J. schenkeli comb. nov. ( Fig. 2 View FIGURES 1–8 ) and J. roeweri comb. nov.; dorsum sparsely covered in short fine setae in males, denser in females; two pairs of weakly sclerotized dorsal sigilla present; ventrally with strongly sclerotised epigyne and booklungs, separated by narrow membranous strips; post-epigastric sclerites broad in males, narrow in females; two paired lines of tiny sclerites between epigastric furrow and spinnerets ( Figs 5, 6 View FIGURES 1–8 ); inframamillary sclerite absent. Genitalia: male palp with large RPA ( Figs 37, 39 View FIGURES 37–40 ), RTA absent or small; palpal femur with shallow longitudinal ventral ridge; palpal patella with lyriform organ at base of RPA on retrolateral side ( Figs 37, 38 View FIGURES 37–40 ); cymbium oval proximally, narrowed distally; tegulum oval, embolus originating retrolaterally or proximally, curving around prolateral margin of tegulum, with tip near distal end of cymbium (e.g. Fig. 39 View FIGURES 37–40 ); conductor and median apophysis absent, except narrow conductor present in J. griswoldi sp. nov. and J. obliquus sp. nov.; female epigyne usually strongly sclerotised, with copulatory openings situated posterolaterally in oblique ( Fig. 40 View FIGURES 37–40 ) or curved ridges, entering directly into large comma-shaped, elongate or oval anterior ST II, with narrow ducts leading to small, posteriorly situated ST I.

Kingdom

Animalia

Phylum

Arthropoda

Class

Arachnida

Order

Araneae

Family

Corinnidae

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