Pseudomacroxiphus C.Willemse 1961

Ingrisch, Sigfrid, 2008, Revision of the genera Paramacroxiphus C. Willemse 1961 and Pseudomacroxiphus C. Willemse 1961 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini), Zootaxa 1755 (1), pp. 1-34 : 30

Pseudomacroxiphus C.Willemse 1961

Type species: Pseudomacroxiphus atrifrons Willemse 1961 , by original designation (= syn of. P. szentia ) Pseudomacroxiphus C. Willemse 1961a , Publ. natuurh. Gen. Limburg 12: 29.

Description. Fastigium verticis narrow, compressed, conical, shorter than scapus; dorsal surface with or without a faint swelling at base; ventral margin separated by a shallow but distinct sinuosity from fastigium frontis. Frons matt or shining, with some shallowly impressed dots; genae subsmooth ( Fig. 82 ). Pronotum subsmooth, rather broad and low, disc broadly rounded into paranota, very apical area slightly raised, flat and shouldered; anterior margin broadly rounded but concave in middle; posterior margin truncate; transverse sulcus faint, broadly interrupted in middle; second transverse sulcus distinct on paranota, obsolete on disc ( Fig. 80 ). Paranota with ventral margin almost straight, descending posteriorly ( Fig. 81 ). Fully winged; tegmen with apex rounded; hind wings of same length as tegmina ( Fig. 81 ). Prosternal spines of medium length (shorter than coxa), but very stout. Mesosternal lobes angular or rounded, metasternal lobes rounded; medial plate at both posterior angles with an obtuse cone. Knee lobes of pro- and mesofemora spinose on both sides (spine on external side of profemur often short, mesofemur on external side sometimes bi-spinose); of postfemur bi-spinose on both sides.

Male. Stridulatory area of tegmen with mirror almost quadrangular and behind mirror with a series of additional fields without nervature, separated from each other by subparallel transverse veins ( Figs. 83–84 ). Tenth abdominal tergite transverse but slightly descending posteriorly; apical margin roundly excised in middle and with a rounded projecting lobe at both sides of excision ( Figs. 90–91 ). Epiproct rounded triangular with a faint medial sulcus in basal half. Projections of paraprocts rather long, stout, conical and curved mediad. Subgenital plate with apex roundly excised in middle, on both sides of excision with short and stout projections which carry the styli; styli as long as or little longer than excised area ( Fig. 91 ).

Female. Tenth abdominal tergite furrowed in midline. Epiproct rounded, surface shallowly depressed. Cerci long, conical, slightly curved, apex acute. Ovipositor long, compressed, blade-shaped, substraight, highest at end of basal third ( Fig. 85 ).

Discussion. Pseudomacroxiphus has the male pronotum not prolonged with the apex truncate and the paranota not deep. A character that it shares with Pseudonicsara and Habetia . It differs from both by the prolonged stridulatory area of the male tegmina and peculiar modifications of the phallic complex. From Pseudonicsara it also differs by the fore tibia which is in cross-section quadrangular and by the long, compressed ovipositor. With Habetia , it also shares the rather flat, not conical meso- and metasternal lobes. It differs, apart from the above characters, by much longer wings, the possession of a baso-internal projection of the male cercus and the male phallic complex. The genus is not related to Macroxiphus . It differs by the morphology of pronotum and sternites, the modified stridulatory apparatus, male abdominal terminalia and titillators, female subgenital plate and even details of the ovipositor, although the latter is prolonged in both genera.

Figures

FIGURES 80–85. Pseudomacroxiphus szentia: 80, male holotype of P. atrifrons; 81, female paratype of P. atrifrons; 82, frons holotype of P. atrifrons; 83, stridulatory area of left tegmen male holotype of P. szentia; 84, do. of right tegmen male paratype of P. atrifrons; 85, ovipositor female paratype of P. atrifrons. Scales = 1 mm (82–84) or 10 mm (81, 85).

FIGURES 86–94. Pseudomacroxiphus szentia details: 86–87, Titillators in ventro-distal view; 88, do. in latero-distal view; 89, do. in dorso-proximal view; 90–91, male abdominal apex in dorsal view; 92 female subgenital plate in ventral view; 93 do. in lateral view; 94, male stridulatory file on underside of left tegmen.—86–87, 91, 94, holotype male of P. szentia; 88–89, holotype male of P. atrifrons; 90, paratype male of P. atrifrons; 92–93, paratype female of P. atrifrons. Abbreviations: 10t tenth abdominal tergite; ce cercus; e epiproct; s stylus; sg subgenital plate. Scales = 1 mm.