Paramacroxiphus elongatus, Ingrisch, 2008

Ingrisch, Sigfrid, 2008, Revision of the genera Paramacroxiphus C. Willemse 1961 and Pseudomacroxiphus C. Willemse 1961 (Orthoptera: Tettigoniidae: Conocephalinae: Agraeciini), Zootaxa 1755 (1), pp. 1-34 : 15-16

Paramacroxiphus elongatus sp. n.

Figs. 14 , 31 , 35 , 48 , 55 , 95

Holotype (male): Indonesia: Papua, North New Guinea Exped., Mamberamo, Alb-Bivak , V.1926, W. Docters v. Leeuwen, labelled: " Pseudonicsara ? elongata sp. n. " . Holotype in Museum Zoologicum Bogoriense [Cibinong], Indonesia .

Description. Fastigium verticis: eye length = 1.5: 2.3 mm; projecting 0.9 mm in front of eyes. Tegmen with anterior margin sinuate: narrowing behind middle, then slightly widening and narrowing again towards apex. Femora with the following number of spines on ventral margins: profemur 7–8 external, 6 internal; mesofemur 6 external, 3–4 internal; postfemur 11 external, 13 internal.

Male. Stridulatory file sinuate, circa 3.26 mm long; teeth large near base, small and dense near apex ( Fig. 14 ); with about 127 teeth or 39.0 teeth per mm; in middle with 31.5 teeth per mm; in basal half with 21.0 teeth per mm. Tenth abdominal tergite with dorsal surface faintly convex in basal area; apical margin prolonged into two triangular lobes, deeply fissing in between; internal apex of triangular lobes slightly curved medioventrad; fissing area, lobes and apex densely setose ( Fig. 31 ). Cerci subcylindrical but ventro-internal surface flattened; dorso-internal margin sub-carinate, prolonged towards apex into a broad-triangular, acute, dorsointernal lobe; apical area bent mediad, compress and at apex divided into a rounded proximal and an acute distal lobe ( Fig. 31 ). Subgenital plate with lateral areas sloping in basal part only; with a faint medial and rounded lateral carinae; central part of subgenital plate with lateral carinae largely prolonged behind and fissing from apex for somewhat less than half the length of the prolongation; styli small ( Fig. 35 ).

Titillators with basal parts simple; apical parts divided into two branches forming semi-membranous sheaths with irregular margins; apico-internal surface with fine striation; marginal parts of lobes darkened and more strongly sclerotised; rim of distal lobe finely crenulated; latero-proximal surface membranous and connected at base with a small, ovoid, membranous sacculus; sheaths of both sides fused medial in basal area ( Figs. 48 , 55 ). Membrane of latero-proximal surface with 2 small, transverse, crenulated lobes. Baso-lateral sclerites large, elongate, tri-carinate.

Female unknown.

Coloration. Brown. Frons with antennal scrobae and mandibles darkened; labrum yellow in central area. Pronotum with a little distinct dark band in dorsal area of paranota. Femora indistinctly mottled with dark and pale areas; meso- and postfemur also with 2 little distinct pre-genicular dark rings. Tegmen maculated, darkened at apex; dorsal area pale.

Measurements of male: body 33; pronotum 10; tegmen 39; postfemur 23 mm.

Diagnosis. P. elongatus belongs to a group of species, in which the central part of the male subgenital plate forms a long apical projection. This character is shared by P. irregularius , P. rufus and P. securiformis . The male cerci resemble the situation in P. rufus although there are distinct differences in shape. P. elongatus however misses the long projections of the tenth abdominal tergite that occur in the latter species. The titillators with weakly sclerotised apical parts that carry a strongly sclerotised cap at tip resembles the situation in P. maculatus , P. rufus and P. securiformis . Details of the structure of the titillators differ however distinctly between all four species.

Etymology. The new species is named for the strongly prolonged male subgenital plate.

Figures

FIGURES 11–15. Stridulatory files on underside of left tegmen of Paramacroxiphus males: 11, P.armatus holotype; 12, P. bifasciatus holotype; 13, P. brunneus holotype; 14, P. elongatus holotype; 15, P. irregularius holotype.

FIGURES 29–34. Male abdominal apex of Paramacroxiphus species in dorsal view (31, in dorso-lateral view; 34, left cercus): 29, P. brunneus holotype; 30, P.armatus holotype; 31, P. elongatus holotype; 32, P. bifasciatus holotype; 33–34, P. irregularius holotype. The arrow points at the baso-internal tooth of the cercus. 10t tenth abdominal tergite; ce cercus; pa process of paraproct; sg subgenital plate.

FIGURES 35–44. Male subgenital plate (35–43) and stridulatory area of right tegmen (44) of Paramacroxiphus species: 35, P. elongatus holotype; 36, P. irregularius holotype; 37, P.rufus holotype; 38, P. securiformis holotype; 39, P.armatus holotype; 40, P. bifasciatus holotype; 41, P. maculatus holotype; 42, P. aberrans holotype; 43, P. brunneus holotype; 44, P.aberransparatype. Abbreviations: m mirror, s scrapper, sp field of spinules.

FIGURES 45–49. Male titillators of Paramacroxiphus species in ventro-distal view: 45, P. brunneus holotype; 46, P. bifasciatus holotype; 47, P. irregularius holotype; 48, P. elongatus holotype; 49, P.armatus holotype (surface structure indicated at apex). Baso-lateral sclerite only shown on one side. Abbreviations: ap apical part of titillator; bp basal part of titillator; br bristles; bs baso-lateral sclerite; m membranous part of phallus.

FIGURES 50–56. Male titillators of Paramacroxiphus species in lateral view (50, 53), dorso-proximal view (51–52, 55– 56) and detail of apical part (54): 50–51, P.armatus holotype; 52, P. bifasciatus holotype; 53–54, P. brunneus holotype; 55, P. elongatus holotype; 56, P. irregularius holotype. Abbreviations: ap apical part of titillator; bp basal part of titillator; br bristles; bs baso-lateral sclerite; m membranous part of phallus. Scale bars = 1 mm.(ZMB); 1 female, do. 67, Pionierlager, 1.v.–30.vi.1912, Bürgers (ZMB); 1 female, do. [label unreadable], Bürgers (ZMB).