Glyptoma schuhi, Irmler, 2015

Irmler, Ulrich, 2015, The neotropical genus Glyptoma ERICHSON, 1839 with descriptions of new species and a key to the species (Coleoptera: Staphylinidae: Osoriinae), Beiträge Zur Entomologie = Contributions to Entomology 65 (2), pp. 297-325 : 318-320

Glyptoma schuhi spec. nov.

( Figs 19 A, B , 23A )

Type material: male Holotype: Peru: Junin Prov., San Ramón de Pangoa , 40 km S Satipo, 750 m elev., 29.1.1974, leg. R. T. Schuh ( AMNH) . Paratypes: 1 male, 4 females with same data as holotype ( AMNH, UIC) ; 3 males, 2 females, Junin Prov. between Cubantia & Masaronquiari, 72 km SE Satipo , 950 m elev., 29.7.1972 ( AMNH, UIC) ; Ecuador: 2 females, Pastaza Prov., Santa Clara env., Puyo , 950 m elev., 18.– 21.7.2008, leg. W. Rossi & I. Tapia ( VAC, UIC) .

Diagnosis: G. schuhi spec. nov. is similarly long as G. obscuricolle ( BERNHAUER, 1908) . It can be easily differentiated from G. obscuricolle ( BERNHAUER, 1908) by the dense setiferous punctation of the elytral disc. In G. schuhi spec. nov. the space between suture and discal carina has at least partly three irregular rows of punctures, whereas in G. obscuricolle only two irregular rows are found.

Description: Length: 3.0– 3.1 mm. Colouration: Black; elytra and antennae dark brown; legs light brown.

Head: 0.45 mm long, 0.60 mm wide; large eyes slightly prominent; longer than temples; temples narrowed to neck in smooth convex curve; sides in front of eyes parallel up to base of antennae; parallel part shorter than length of eyes; at base of antennae with obtuse nearly rectangular angle; narrowed to anterior edge of clypeus in sinuate curve; anterior edge of clypeus less wide than half of width between base of antennae; setiferous punctation moderately dense and deep; on average, interstices between punctures less wide than half of diameter of punctures; setae nearly twice as long as interstices between punctures; without microsculpture; surface polished.

Antenna distinctly longer than head and pronotum combined; all antennomeres nearly equally wide; second antennomere short; nearly twice as wide as long; tenth antennomere only one third wider than long; third to tenth antennomere with numerous long yellow setae; setae approximately as long as width of antennomeres.

Pronotum: 0.52 mm long, 0.55 mm wide; widest in anterior third; narrowed to anterior widely rounded angles in smooth convex curve; narrowed to posterior obtuse angles in concave curve; setiferous punctation deep and dense; width of punctures irregular; interstices between punctures restricted to small ridges; insertion of setae partly asymmetric; at posterior angles, punctation partly coriaceous; without micro¬sculpture; surface of ridges polished.

Elytra: 0.79 mm long, 0.78 mm wide; widest in posterior third, but nearly parallel; shoulders shortly rounded to emarginate anterior edge; discal carina irregular; partly broken; lateral carina absent; suture line marked by small coriaceous punctures; not continued to posterior margin; at posterior margin punctures larger; between suture line and discal carina two to three longitudinal rows of setiferous punctures; row of punctures adjacent to suture line and at inner margin of discal carina complete; intermediate row restricted to few irregular punctures; lateral row denser than inner row; laterally to discal carinae several rows of setiferous punctures; punctures of these rows larger than punctures of inner rows and partly coriaceous; interstices mostly restricted to small ridges; without microsculpture; ridges polished.

Abdomen with dense and large setiferous punctation; interstices between punctures restricted to extremely small ridges; surface with deep and wide netlike microsculpture; microsculpture on posterior segments much weaker; anterior segments matt; posterior two segments shiny.

Aedeagus smoothly curved from basal to apical lobe; apex acute; apical lobe laterally with numerous sensillae; apical sensillae smaller than basal sensillae; paramere short; reaching basal fourth of apical lobe; slightly sinuate; basal half thick; in anterior half abruptly thinner.

Etymology: The species is named to honour the collector R. T. Schuh, who collected intensively in the Neotropical region.

Key to the species

1 Large species of 3.5 to 4.0 mm, antennomeres longer than wide or quadrate ............................................................ 2

- Species smaller than 3.5 mm (max. 3.2 mm), antennomeres wider than long ........................................................... 6

2 Head without any microsculpture, totally shiny ..................................................................... G. laeviceps SHARP, 1887

- Head at least partly at supraocular areas with microstriae, partly or totally matt or with iridescent shine ............ 3

3 Head, pronotum and elytra with dense isodiametric microsculpture, totally matt ......... G. opacum ( SHARP, 1876)

- At least clypeus shiny and without microsculpture ........................................................................................................ 4

4 Elytra with discal and at least weak lateral carina ........................................................................................................... 5

- Elytra only without discal carina ............................................................................. G. cicatricosa MOTSCHULSKY, 1857

5 Head with supraocular fine striate microsculpture, with iridescent shine ...................................... G. rossi spec. nov.

- Head except clypeus with dense isodiametric microsculpture and matt ..................................................................... 6

6 Pronotum with large pseudo-punctures; setiferous punctures mostly on ridges between pseudo-punctures ....... 7

- Pronotum with normal setiferous punctation, interstices between punctures wider .............................................. 12

7 Elytra wider than long, pseudo-punctures of elytra combined to larger deep impressions ......................................... .................................................................................................................................... G. guadelupense (WENDELER, 1930)

- Elytra quadrate or slightly longer than wide, pseudo-punctures of elytra isolated or combined to smaller grooves ... 8

8 Small species of 2.3 mm length, head without microsculpture, surface shiny, discal carinae of elytra distinctly convergent to scutellum ........................................................................................................... G. hirticorne SHARP, 1887

- Larger species of at least 2.5 mm length, head with dense microsculpture and matt, discal carinae of elytra not distinctly convergent to scutellum, slightly curved ........................................................................................................ 9

9 Elytra slightly longer than wide, pseudo-punctures of pronotum and elytra isolated ............................................. 10

- Elytra quadrate, pseudo-punctures of elytra at least partly combined to small grooves, pseudo-punctures of pronotum partly combined to irregular deep grooves ............................................................................................................ 11

10 Head with weak microsculpture, surface shiny ............................................................ G. crassicorne ERICHSON, 1840

- Head with dense and deep microsculpture, surface matt ............................................................ G. lescheni spec. nov.

11 Temples convergent to neck, slightly convex, pseudo-punctures on pronotum less deep .... G. guianense spec. nov.

- Temples angulated, pseudo-punctures of pronotum partly combined to large deep grooves ..................................... ...................................................................................................................................................... G. amazonense spec. nov.

12 Small species of less than 1.9 mm length, elytra distinctly longer than wide, L/W at least 1.2 ................................... ........................................................................................................................................... G. klimschi ( BERNHAUER, 1908)

- Larger species of at least 1.9 mm length, elytra quadrate or only slightly longer than wide, L/W not more than 1.1 .............................................................................................................................................................................. 13

13 Space between suture and discal carina with at least two irregular rows of punctures in posterior half .............. 14

- Space between suture and discal carina with impunctate areas and only one row in posterior half ..................... 15

14 Large species of at least 3.0 mm length, space between suture and discal carina with dense punctation, partly with more than two parallel rows .............................................................................................................. G. schuhi spec. nov.

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Smaller species of less than 3.0 mm length, space between suture and discal carina with two parallel irregular rows of punctures ........................................................................................................................ G. nitens ( BERNHAUER, 1908) 15 Larger species of more than 3.0 mm length, vertex of head weakly punctate and with micro-striae, surface with slight iridescent shine .............................................................................................. G. obscuricolle ( BERNHAUER, 1908)

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Smaller species of 1.9–2.4 mm length, punctures on vertex of head larger .............................................................. 16 16 Elytra without microsculpture, shiny ........................................................................... G. nitidum ( BERNHAUER, 1908)

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Elytra on impunctate space with fine longitudinally striate microsculpture, with iridescent shine or matt ........ 17 17 Slightly larger species of 2.4 mm length, elytra slightly longer than wide, microsculpture denser and on impunctate space of elytra with iridescent shine .............................................................................. G. denieri ( BERNHAUER, 1939)

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Slightly smaller species of 2.1–2.2 mm length, elytra quadrate, microsculpture weaker and impunctate space of elytra shinier .............................................................................................................................. G. cognatum SHARP, 1887

New combination

Figures

Figs 23: Fore body of Glyptoma

Figs 18–19: Aedeagus (A) and antenna (B) of Glyptoma klimschi (18), G. schuhi (19); scale bar: 0.1 mm.