Therates motoensis Tan, 1981

Matalin, Andrey V. & Wiesner, Jürgen, 2023, Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae), Zootaxa 5256 (5), pp. 401-433 : 421

Therates motoensis Tan, 1981

Figs 13, 14 , 40, 41 , 62, 63 , 97, 98 , 172 .

Therates motoensis Tan, 1981: 332 (Type locality— Xizang, Mêdog).

Therates fruhstorferi vitalisi W. Horn, 1913 — Wiesner 1988: 40, 1992: 92; Lorenz 1998: 38, 2005: 117; Puchkov & Matalin 2003: 115.

Therates motoensis Tan, 1981 — Wu 2011: 31, 37; Wiesner & Bi 2014: 233; Putchkov & Matalin 2017: 245; Wiesner 2020: 95.

Material. 1♂ 1♀ — China, Xizang, Motuo, Hanmi , 2100 m, 15–30.VII.2013, leg. Wen-Xuan Bi ( JW) .

References. 3♂♂ 3♀♀ — China, Xizang, Motuo, Hanmi , 2100 m, 15–30.VII.2013, leg. Wen-Xuan Bi ; 19♂♂ 17♀♀ — ibid, 12–24.VII.2013 ; 6♂♂ 8♀♀ — ibid, 25–31. VII.2013 ; 6♂♂ 2♀♀ — ibid, 23–28.VII.2011 ; 2♂♂— China, Xizang, Motuo , 1700–2000 m, 18.VIII.2013, leg. Wen-Xuan Bi ; 2♂♂ 1♀ — China, Xizang, Motuo , 2100 m, 3.VIII.2012, leg. Chao Wu ; 1♀ — China, Xizang, Motuo , Hanmi-A’niqiao, VIII.2005, leg. Hao Huang ( Wiesner & Bi 2014).

Diagnosis. From all related species T. motoensis is readily distinguished by the bluish-green topside ( Figs 13, 14 ); by the colour of labrum ( Figs 62, 63 ); as well as by the aedeagus clearly dilated in apical third ( Fig. 172 ).

Redescription. TL = 11.2–12.9 mm (mean = 12.1 mm, n = 6).

Head shining bluish-green with light golden reflection; orbital plates striated throughout or finely striated in posterior third; frons convex, semi-circular, smooth or slightly rugose, frontal sulci deep especially in anterior third, convergent in anterior 2/3, shallow and distinctly divergent in posterior third. Mandibles light brown underside, pale-yellow topside with light-brown teeth and apical molar. Labial and maxillary palpi yellow with slightly darker basal palpomeres. Antennae lanceolate, extending posteriorly nearly to the elytral shoulders in males, somewhat shorter in the females, scape yellow or yellow-brown on anterior side and brown-black on posterior side, remaining antennomeres black, antennomeres 2–5 with light blue tinge, antennomeres 6–11 approximately equal width without protruding downward anterior lower margins ( Figs 40, 41 ). Labrum indistinctly transverse, LW/LL = 1.0–1.14 (mean = 1.07, n = 4), with six–eight apical teeth and seven–nine setae between them, yellowish with black margins and large basal spot in males ( Fig. 62 ), black with yellowish oval apical spot in females ( Fig. 63 ).

Pronotum slightly transverse, PW/PL = 1.0–1.1 (mean = 1.03, n = 4), shining bluish-green with light golden tinge, apical lobe slightly wider in middle portion; thorax greenish-blue with light golden tinge.

Fore and middle femora pale-yellow along posterior side and black-brown with bluish tinge along anterior side, hind femora pale-yellow in basal half and black with blue tinge in apical half; fore and middle tibiae pale-yellow except narrow brown-black apices, hind tibiae pale-yellow except brown-black basal third, distinctly wider than fore and middle tibiae; tarsomeres 1 and 2 of fore- and middle legs pale-yellow or light brown with darker apices, other tarsomeres dark-brown, all hind tarsomeres pale-yellow except dark-brown apices; HTbL/HTaL = 1.09–1.15 (mean = 1.12, n = 2).

Elytra prolonged, indistinctly divergent toward apex, EL/EW = 2.04–2.13 (mean = 2.08, n = 4); longer in females—EL/PL = 4.0–4.25 (mean = 4.13, n = 2) than in males—3.7–3.8 (mean = 3.75, n = 2); shining green with golden reflection; punctuation deep and regular between shoulders and central dot, shallower behind, in apical quarter virtually indistinct; scutellum dark blue with golden-green tinge; apical margin gradually rounded. Elytral pattern presented by very small invisible dorsally yellow basal portion of humeral lunule and small trapezoid yellow central dot ( Figs 13, 14 , 97, 98 ).

Aedeagus clearly dilated in apical third, with gradually sinuate, prolonged apex ( Fig. 172 ), AL = 2.75–2.9 mm (mean = 2.8, n = 2), EL/AL = 2.69.

Distribution. CHINA: Xizang (Mêdog County) ( Fig. 184 ).

Figures

FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png.

FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels.

FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales.

FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes.

FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype.

FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm.