Therates pseudovitalisi tibetana Matalin et Wiesner, 2023

Matalin, Andrey V. & Wiesner, Jürgen, 2023, Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae), Zootaxa 5256 (5), pp. 401-433 : 408-409

Therates pseudovitalisi tibetana Matalin et Wiesner , ssp. n.

Figs 11 , 38, 39 , 60, 61 , 94, 95 , 135 .

Therates fruhstorferi vitalisi W. Horn, 1913 — Sawada &Wiesner 2006: 463; Wu 2011: 31.

Type material. HOLOTYPE, 1♂— China, Tibet, Cha Yu [= Zayü] County , Xia-Cha-Yu, 1900–2600 m, 1– 28.7.2004 ( JW) ; PARATYPE, 1♀ —same labeled as the holotype ( JW) .

Diagnosis. This new subspecies is best distinguished from the nominotypical form by the shape of antennomeres 9–11 ( Figs 38, 39 vs. Figs 36, 37 ); by the proportions of hind tibiae and tarsi—mean HTbL/HTaL = 1.25 vs. HTbL/ HTaL = 1.32; by virtually transverse, clearly dilated in lateral portion central dot ( Figs 11 , 94, 95 vs. Figs 9, 10 , 90–93 ) and by shorter elytra—mean EL/AL = 2.48 vs. mean EL/AL = 2.61.

Description. TL = 10.2 mm in male, 11.6 mm in female.

Head shining black-blue with light violet reflection; orbital plates slightly striated in posterior third; frons semi-circular (see above), slightly downward anteriorly (see at the side), smooth, frontal sulci deep and indistinctly convergent in anterior 2/3, shallow and sharply divergent in posterior third; occiput flat or slightly concave in anterior margin. Mandibles in male brown underside, pale-yellow topside with brown teeth and apical molar; in female dark brown except yellow-brown basal half of topside. Two basal labial palpomeres light brown, palpomere 3 pale-yellow with brown apex, apical palpomere dark brown; palpomeres 1 and 2 of maxillary palpi brown, palpomeres 3 and 4 brown-black. Antennae barely extend shoulders, antennomeres 9–11 in males distinctly dilated and slightly club-shaped, antennomeres 9 and 10 with protruding downward anterior lower margins, antennomere 11 with sharply cut straight anterior margin ( Fig. 39 ); antennomeres 9 and 10 in female only slightly protruding downward, antennomere 11 oval with protruding apex ( Fig. 38 ). Labrum as long as width (LL = LW— 1.2 mm in male; 1.5 mm in female), in male black-brown with medium-sized yellow-brown apical spot clearly separated from central apical teeth ( Fig. 60 ), in female with small barely visible brown spot in anterior third ( Fig. 61 ).

Pronotum shining black-blue; indistinctly transverse, PW/PL = 1.05–1.06 (mean = 1.05, n = 2), apical lobe indistinctly wider in central portion; thorax shining blue with light golden reflection.

Fore femora yellow-brown with blue-tinged posterior side; middle femora in male black-brown with yellowbrown base and light blue-tinged anterior margin, in female entirely brown with blue tinge; hind femora pale-yellow in basal third and black with blue tinge in apical 2/3; fore and middle tibiae in male light brown in basal third and dark-brown blue-tinged in apical third; hind tibia entirely black with violet reflection; in female all tibiae black with blue lustre; tarsi black with indistinct blue tinge; HTbL/HTaL = 1.24–1.25 (mean = 1.25, n = 2).

Elytra shining blue with light purple-violet reflection; indistinctly divergent toward apex, EL/EW = 1.95–1.97 (mean = 1.96, n = 2); punctuation deep and regular between base of humeral hump and central dot, shallower and sparse in the middle and practically absent in apical quarter; scutellum black with blue tinge; apical margin clearly cut. Elytral pattern consists of pale-yellow very small basal portion of humeral lunule (see in front) and elongate practically transverse central dot clearly dilated in lateral portion; basal dot virtually invisible in male, entirely absent in female ( Figs 11 , 94, 95 ).

Aedeagus gradually sinuate with elongate rounded apex lacking lateral carinae ( Fig. 135 ); AL = 2.6 mm, EL/AL = 2.48.

Etymology. This new species is named after the type area—Chinese Tibet Autonomous Region.

Distribution. CHINA: Tibet Autonomous Region (Cha Yu [= Zayü] County) ( Fig. 184 ).

Figures

FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png.

FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels.

FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales.

FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes.

FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype.

FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm.