Tingupa Chamberlin, 1910

Shear, William A. & Shelley, Rowland M., 2007, Tingupa tlingitorum, n. sp., a new milliped from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida: Tingupidae), Zootaxa 1393, pp. 53-59 : 55

Genus Tingupa Chamberlin, 1910

Tingupa Chamberlin, 1910:238 . Chamberlin & Hoffman, 1958:106. Buckett, 1964:14. Shear, 1969:141; 1972:265; 1981:6­7. Jeekel, 1971:82. Hoffman, 1980:135; 1999:254­255. Kevan, 1983:2967. Shelley et al., 2000:79. Type­species. T. utahensis Chamberlin, 1910 , by original designation.

Diagnosis. Tingupidae with prominent, rounded paranota, metaterga with microsculpture of low tubercles interspersed among short, sharp ridges (see Shear 1981, figs. 1–2).

Distribution. Same as that of the family but excluding Colorado, West Virginia, Virginia, and North Carolina ( Fig. 4 ). Idaho and the Queen Charlotte Islands are based on the following samples of females that are unidentifiable to species; the latter site is some 450 mi (720 km) south­southwest of Haines.

USA: IDAHO, Valley Co., 4 mi (6.4 km) NE McCall, Ψ, 18 October 1944, W. Ivie (NMNH). New State Record.

CANADA: BRITISH COLUMBIA : QUEEN CHARLOTTE ISLANDS, Graham I., Queen Charlotte City, 3Ψ, 9 October 1948, G. J. Spencer (RBCM). New Country and Provincial Record.

Remarks. Tingupa comprises ten species, plus the new one described herein, but only nine are keyed in Shear (1981) and cited by Hoffman (1999), who both missed T. intergerina Loomis & Schmitt, 1971 , in western Montana. Six species inhabit the northwestern Pacific Coast in the "lower 48" from east­central California to northwestern Oregon; one species occurs in western Montana; the type­species, ostensibly with two subspecies, occupies the Wasatch Mountains and Parowan Valley of Utah; another nominal species, probably a synonym of T. utahensis , inhabits the Santa Catalina, Pinaleno , and Chiricahua Mountains of southeastern Arizona; and a troglobitic species occupies limestone caves in southwestern Illinois, Missouri, and northern Arkansas ( Chamberlin 1910, 1925, 1928; Chamberlin & Hoffman 1958; Loomis & Schmitt 1971; Shear 1972, 1981; Hoffman 1999).

Figures

FIGURES 2 – 4. Tingupa tlingitorum. 1, gonopods, anterior view. 2, colpocoxite branches, posterior view. 3, right leg 9, anterior view.