Physodera bifenestrata Heller, 1923

Ma, Yunlong, Shi, Hongliang & Liang, Hongbin, 2017, Revision of the Oriental Genus Physodera Eschscholtz, 1829 (Coleoptera, Carabidae, Lebiini, Physoderina), with the descriptions of two new species, Zootaxa 4243 (2), pp. 297-328 : 304-305

Physodera bifenestrata Heller, 1923

Fig. 18 , 39 .

Heller 1923: 304 (type locality: Sandakan (Borneo); lectotype in SNSD); Csiki 1932: 1347; Jedlička 1963: 301; Shi et al. 2013: 41.

Material examined. Lectotype of Physodera bifenestrata Heller , designated herein (SNSD), a male, "Sandakan / Borneo / baker"; "1920"; " bifenestrata / Typus" [red label]; "Staatl. Museum für / Tierkunde, Dresden". ( Figs. 6 , 18 ); 1 male (MNHN), "Borneo Occ., Pontianak, 1900"; " Physodera spec."; "Ritsema, vidit 1901."; "Museum Paris, 1952, Coll. R. Oberthür".

Notes on type. In the original literature, Heller mentioned the locality "Sandakan, Borneo", but didn't indicate the number of type series. We borrowed a male from SNSD bearing a type label and perfectly fitting with the type locality. We herein designate this specimen as lectotype for the purpose of fixing the name to unique name-bearing type.

Diagnosis. Body length 9.1–9.9 mm. Dorsal side black with metallic purple hue; pronotum unicolor ; elytra with a pair of ivory callosities on disc, callosity about 0.4–0.5 times as elytron in length, placed in the middle of elytron, ovate in shape, without notch. Tergum and sternum VII without distinct pattern. Antennameres 5–11 distinctly widened and flattened (Fig. 59). Elytral third interval with two setigerous pores, the fifth with only one pore near base; apical margin truncated.

This species can be distinguished from related species ( P. eburata and P. sciakyi ) by: elytral callosities larger, more than 0.4 times as the elytra length; pronotum wider, greatest width about 1.9 times as the median length (PW/ PL less than 1.7 in the other two species).

Male genitalia ( Fig. 39 ). Median lobe of aedeagus fairly stout and straight, apex barely bent to right side in ventral view, right margin weakly angulated before apex; apical lamella very small, its length about half as basal width, about triangular, rounded apically. Internal sac with main flagellum thick and straight, apex almost reaching apical orifice; secondary flagellum short and thick, weakly sclerotized, length about 0.3 times as the main flagellum; trumpet-form expansion large, length about 0.4 times as the main flagellum; apical bursa present.

Female genitalia. Unknown.

Distribution. Only known from Borneo. ( Fig. 65 )

Figures

FIGURE 39. Median lobe of male genitalia, right-lateral, ventral, left-lateral, and dorsal view of P. bifenestrata Heller, Borneo, scale bar = 0.5 mm.

FIGURE 65. Known distribution of Physodera eburata group: * P. eburata Heller; ● P. bifenestrata Heller; Ɨ P. sciakyi sp. n., type localities for valid names in red.

FIGURES 13 – 18. Habitus of type materials, scale bars = 2 mm: Fig. 13, holotype of P. diglena Andrewes; Fig. 14, syntype of P. dejeani Eschscholtz; Fig. 15, syntype of P. noctiluca Mohnike; Fig. 16, holotype of P. chalceres Andrewes; Fig. 17, lectotype of P. eburata Heller; Fig. 18, lectotype of P. bifnestrata Heller.

FIGURES 1 – 12. Labels for type materials: Fig. 1, holotype of P. diglena Andrewes; Fig. 2, syntype of P. dejeani Eschscholtz; Fig. 3, syntype of P. noctiluca Mohnike; Fig. 4, holotype of P. chalceres Andrewes; Fig. 5, lectotype of P. eburata Heller; Fig. 6, lectotype of P. bifnestrata Heller; Fig. 7, holotype of P. sciakyi sp. n. Fig. 8, holotype of Lachnoderma andrewesi Jedlička; Fig. 9, holotype of P. bacchusi Darlington; Fig. 10, holotype of P. unicolor, sp. n.; Fig. 11, lectotype of P. eschscholtzii Dejean; Fig. 12, lectotype of P. davidis Fairmaire.