Geodromicus (s.str.) kashmire nsis Cameron, 1930

Shavrin, Alexey V., 2022, The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae), Zootaxa 5213 (5), pp. 451-496 : 472-474

Geodromicus (s.str.) kashmire nsis Cameron, 1930

( Figs. 40–43 , 46–51 )

Geodromicus kashmirensis Cameron, 1930: 455

Geodromicus similis Cameron, 1930: 455 syn.n.

Geodromicus consors Cameron, 1945: 65 syn.n.

Geodromicus (Geodromicus s.str.) afghanicus Scheerpeltz, 1961: 36 syn.n.

Geodromicus afghanensis Coiffait [sic!]: Tronquet 1981: 69

Geodromicus karakorus Coiffait, 1977: 269 syn.n.

Type material examined. Lectotype (here designated) of Geodromicus kashmirensis Cameron, 1930 , ♂ ( Fig. 20 ; dissected): ‘Type’ <round label with red margin, printed>, ‘Ferozepur Nala 6500 | Gulmarg, Kashmir. | B. M. Bhatia. | 2.[handwritten] VI. 1928.’ <printed>, ‘Under | wet stones’ <printed>, ‘311’ <handwritten>, ‘M.Cameron | Bequest. | B.M.1955-147.’ <printed>, ‘G. | kashmiricus | TYPE Cam’ <handwritten>, ‘ LECTOTYPE | Geodromicus | kashmirensis Cameron, 1930 | Shavrin A.V. des. 2022’<red, printed>, ‘ Geodromicus (s.str.) | kashmirensis Cameron, 1930 | Shavrin A.V. det. 2022’ <printed> ( BMNH).

Lectotype (here designated) of Geodromicus similis Cameron, 1930 , ♀ ( Fig. 41 ; right antennomeres 9–11 glued on the same card with the specimen): ‘Type’ <round label with red margin, printed>, ‘Ferozepur Nala 6500 | Gulmarg, Kashmir. B.M. Bhatia. | 2. [handwritten, in black] VI. 1928.’ <printed>, ‘Under | wet stones’ <printed>, ‘313 <handwritten>, ‘ Geodromicus | similis | TYPE [in red, underlined] Cam.’ <handwritten>, ‘M. Cameron. | Bequest. | B.M. 1955-147.’ <printed>, ‘ LECTOTYPE | Geodromicus | similis Cameron, 1930 | Shavrin A.V. des. 2022’ <red, printed>, ‘ Geodromicus (s.str.) | kashmirensis Cameron, 1930 | Shavrin A.V. det. 2022’ <printed> ( BMNH).

Holotype of Geodromicus consors Cameron, 1930 , ♀ ( Fig. 42 ): ‘Type’ <round label with red margin, printed>, ‘Kashmir | Gulmarg | vi-vii-31 | Dr. Cameron’ <printed>, ‘M. Cameron. | Bequest. | B.M. 1955-147.’ <printed>, ‘G. | consors | TYPE [red, underlined] Cam.’ <handwritten>, Geodromicus (s.str.) | kashmirensis Cameron, 1930 | Shavrin A.V. det. 2022’ <printed> ( BMNH).

Holotype of Geodromicus (Geodromicus s.str.) afghanicus Scheerpeltz, 1961 , ♀: ‘Panjao 24 [handwritten] / 7 [handwritten] 1948 | Afghanistan N. Haarløv | St. 85. [handwritten]’ <printed>, ‘3. Danske Exp. III | Centralasien’ <printed>, ‘ ♀ ’ <handwritten>, ‘TYPUS | Geodromicus [handwritten] | afghanicus [handwritten] | O.Scheerpeltz.’ <red, printed>, ‘ Geodromicus | afghanicus | nov. spec. | det. Scheerpeltz, 19 [printed] 58’ <handwritten>, ‘ZMUC | 00523323’ <printed>, ‘ Holotypus ♀ | Geodromicus | afghanicus | Scheerpeltz | V.Gusarov rev. 1993’ <red, handwritten>, ‘ Geodromicus ♀ | afghanicus Scheerp. | V.I. Gusarov det. 1993 [printed]’ <handwritten> ( ZMUC).

Holotype of Geodromicus karakorus Coiffait, 1977 , ♂ ( Fig. 43 ; dissected): ‘Nangmah-Tal | Karakorum, 1.8.[19]70 | lg. Koblmüller’ <handwritten>, ‘ Geodromicus | spec. | det.H.Franz [printed]’ <handwritten>, ‘TYPE’ <red, printed>, ‘ Geodromicus | karakorus | H. COIFFAIT det. 19[printed]76’ <handwritten>, ‘ Geodromicus | kashmirensis Cameron, 1930 | Shavrin A.V. det. 2022’ <printed> ( NMW).

Additional material examined. AFGHANISTAN: 1 ♂: ‘Kot Gai 2500m 21.VIII.[19]74’, ‘Palktia Afghan[istan].’, ‘Ft. de C...[unreadable] G.M[eurgues]. et G.L[edoux] ’, ‘MUSEUM PARIS | COLL. J. JARRIGE’, ‘ Geodromicus afghanensis Coiffait’ ( MHNH); 1 ♀: ‘St. 280’, ‘Voyage en Afghanistan K. Lindberg’, ‘Bend-Amir 1.8.[19]57’, ‘…[unreadable] Steien.’ ‘ Geodromicus afghanicus Scheerp. ’, ‘coll. Scheerpeltz’ ( NHMW); 1 ♂: ‘M. Daniel lgt by a trap on a lorry’, ‘ Kabul —12.VI. Tangi Gar [19]67. AFGHANISTAN’ ( NMPC); 1 ♂: ‘COTYPUS Geodromicus afghanicus O. Scheerpeltz’, ‘ ♂ ’, ‘J. Klapperich Pagmangebirge 2300 m, 30.5.[19]52 O. Afghanistan’, ‘MUSEUM KOENIG BONN’, ‘ Geodromicus afghanicus H.COIFFAIT det. 1981’( MHNH); 1 ♀: ‘ ♀ ’, ‘Afghanistan Badagsthan’, ‘COTYPUS Geodromcius afghanicus O. Scheerpeltz’, ‘ Geodromicus afghanicus | 3278B n.sp.’, ‘Collection Arthur Spälti’( MHNG); 1 ♂ [additional printed label: ‘m’], 1 [additional printed label: ‘f’]: ‘Paratypus’, ‘COTYPUS Geodromicus afghanicus O. Scheerpeltz’, ‘J. Klapperich Pagmangebirge [Pagman] 2400 m, 6. 7. [19]52 O. Afghanistan’, ‘H. Coiffait det. 1981’, ‘MUSEUM KOENIG BONN’, ‘ Geodromicus afghanicus Scheerp.,1960 V.I. Gusarov det 1993’ ( ZFMK); PAKISTAN: 1 ♂, 1 ♀: Khyber Pakhtunkhwa, Great Karakorum Mts., to west from Chikar. 3200–3500 m a.s.l.. 01– 10.08.2009. V. Gurko leg. (cSch); 1 ♂: Gilgit-Baltistan, Hunza-Nagar District, W Chikar. 3200–3500 m a.s.l. 01– 10.08.2009.V.Gurko leg. (cSh); INDIA: KASHMIR: 13 ♂♂, 24 ♀♀, 1 spec.: ‘Kashmir Gulmarg vi-vii-31 Dr. Cameron’, ‘M. Cameron. Bequest. B.M. 1955-147.’ ( BMNH); 3 ♀♀ (one specimen with additional label: ‘consors Cam’): ‘Paratype’, ‘Kashmir Gulmarg vi-vii-31 Dr. Cameron’, ‘M. Cameron. Bequest. B.M. 1955-147.’ ( BMNH); 2 ♂♂, 2 ♀♀: ‘Gulmarg Kashmir 6/[19]31. Cameron’, ‘ similis Cam. auct. det.’, ‘W. Steel coll. B.M. 1969-552’; 1 ♂, 4 ♀♀: ‘Kashmir Gulmarg 8,000 –9,000 ft. vi-vii-1931 Dr.Cameron’, ‘W. Steel coll. B.M. 1969-552’ ( BMNH); 2 ♀♀: ‘Kashmir Khelanmarg 10,000 ft. vi-vii-1931 Dr. Cameron’ ( BMNH); 1 ♀: ‘Kashmir Lianmarg 10,000 ft. vi-vii-1931 Dr.Cameron’ ( BMNH); 2 ♂♂, 1 ♀, 3 exs.: Tangmarg Pir Panjal. 2600 m a.s.l. 21– 25.05.1976. Martens & Schawaller leg. ( MHNG, MHNH); 2 ♂♂: ‘Kashmir’ ( BMNH); LADAKH: 1 ♂, 1 ♀: Chemre. 06.1995. Sauer leg. (cSch); 3 ♂♂ (specimens from FMNH with additional label: ‘amnicola Cam. det. Bernhauer Standinger don.’), 4 ♀♀: ‘Gya-Ladak Taglang-Pass Himalaya mont.’ ( FMNH, NMPC, SNSD).

Redescription. Measurements (n=50): HW: 0.90–1.18; HL: 0.50–0.79; OL: 0.26–0.30; TL: 0.10–0.15; AL (averaged): 3.42; PL: 0.80–1.45; PWmax: 1.07–1.60; PWmin: 0.72–1.10; ESL: 1.71–2.25; EW: 1.82–2.25; MTbL (averaged): 1.55; MTrL (averaged): 0.60 (MTrL 1–4: 0.40; MTrL 5: 0.20); AW: 1.70–2.10; AedL: 1.05–2.05; BL: 4.25–7.20 (holotype of G. kashmirensis : 7.00; holotype of G. karakorus : 5.20; holotype of G. consors : 6.80; lectotype of G. similis : 5.00).

Habitus as in Figs. 40–43 . Body reddish-brown to dark-brown, sometimes with darker head, paler middle portion of elytra and yellowish intersegmental membranes of abdomen and/or paratergites in some specimens; mouthparts, antennae and legs yellow-brown to brown (some specimens with darker antennomeres 1–2 or 1–3 and tibiae). Head without or with distinct shallow and transverse microreticulation in middle; neck with very dense rugose and isodiametric microsculpture; pronotum without or with very indistinct transverse meshes in medioapical portion.

Head 1.4–1.8 times as broad as long, with slightly or distinctly elevated infraorbital portions, frontal portion with relatively strongly convex supra-antennal elevations, with very deep, wide and subtriangular anterio-median depression, temples wide and convex, or distinctly narrowed posteriad behind posterior margins of eyes, about twice as long as longitudinal length of eyes; interocellar depression variable, deep and wide, or slightly impressed and narrow, separated from infraorbital portions by moderately deep, long anteocellar foveae (indistinct in some specimens), reaching level of middle or apical third of eyes. Ocelli small or moderately large, distance between ocelli 1.3–1.7 times as long as distance between ocellus and posterior margin of eye. Punctation irregular, moderately sparse and fine: denser, slightly larger and deeper in middle and infraorbital portion, or finer and sparser in middle and larger and denser on infraorbital portions. Antenna reaching middle length or apical third of elytra when reclined; antennomere 3 1.3–1.4 times as long as 2, antennomere 4 slightly broader and distinctly shorter than 3, antennomeres 5–10 indistinctly longer than 4 (sometimes antennomeres 8–10 slightly broader than 7).

Pronotum 1.3–1.4 times as broad as long, from widest middle or medio-apical third strongly narrowed toward moderately long subparallel laterobasal margins, with shallow or moderately deep transverse mediobasal depression. Punctation dense, larger and deeper than that in middle portion of head, sometimes finer and sparser in mediobasal portion (some specimens with small oval impunctate portion in front of mediobasal depression).

Elytra 1.4 times to twice as broad as long, slightly or widely broadened posteriad, 1.5 times to twice as long as pronotum. Punctation dense, about as that on pronotum or slightly larger and deeper, denser and finer in parascutellar portion and sometimes sparser and finer along suture.

Metatibia more than twice as long as metatarsus.

Abdomen distinctly narrower than elytra, with two very large and transverse tomentose spots in middle of abdominal tergite IV.

Male. Pronotum large, very wide and convex. Profemuri and protarsomeres 1–4 wide. Apical margin of abdominal tergite VIII straight or slightly concave. Apical margin of abdominal sternite VIII concave. Aedagus with moderately wide basal part, gradually narrowed toward rounded or subacute apex; aedeagus widely broadened apicad, reaching or slightly exceeding apex of median lobe, with four moderately short apical setae; internal sac with very narrow and moderately short flagellum in basal or middle portion ( Figs. 46–49 ). Lateral aspect of the aedeagus as in Fig. 50 .

Female. Pronotum small, moderately narrow, less convex. Profemuri and protarsomeres 1–4 narrow. Apical margins of abdominal tergite VII and sternite VIII rounded.

Comparative notes. Based on the coloration, body length and the general shape of the body, G. (s.str.) kashmirensis is similar to G. (s.str.) convexicollis , from which it can be distinguished by the less transverse pronotum, distinctly narrower median lobe and broader apical portions of the parameres.

Distribution. Geodromicus (s.str.) kashmirensis is known from several localities of Afghanistan, Pakistan and India (Ladakh, Kashmir) ( Fig. 51 ).

Bionomics. Specimens were collected at elevations from 1900 to 3500 m a.s.l. and were taken from under stones near streams and rivers.

Remarks. Geodromicus kashmirensis was originally described based on unspecified number of syntypes from “Gulmarg, Ferozepur Nala, alt. 6500 feet …”. I designate here the studied specimen from BMNH as the lectotype in order to fix the ideltity of the name.

Geodromicus similis was originally described based on unspecified number of syntypes from the same locality as that in G. kashmirensis . I designate here this specimen (female) as the lectotype in order to fix the identity of the name. Based on the morphological features, it conspecific with the lectotype of G. kashmirensis and other studied specimens of this species. Thus, I synonymized G. similis with G. (s.str.) kashmirensis .

Geodromicus consors was originally described based on the holotype (“Type in my collection”) from “Kashmir: Gulmarg”. I studied this type and found that it is conspecific with the lectotype of G. kashmirensis and other studied specimens of G. (s.str.) kashmirensis . Consequently, G. consors is synonymized with it.

Geodromicus (Geodromicus s.str.) afghanicus was described based on the female from “Panjao in Afghanistan...”. Tronquet (1981) cited it as “ G. afghanensis Coiffait (sous presse)” from “Khot Gai, massif du Tarte Soleiman, au Sud de Djalalabad, 2 500 m …” and provided figures of the habitus ( Tronquet 1981: fig. 3) and the aedeagus ( Tronquet 1981: figs. 4: A–B). Based on the study of the holotype and additional material, I found that G. afghanicus is morphologically conspecific with G. (s.str.) kashmirensis . In consequence, G. afghanicus is synonymized with it.

Geodromicus karakorus was originally described based on the holotype and paratype from “Vallée Nang Mah, Karakorum…”. Based on the study of the holotype, I found that this species based on the external and internal morphological features is conspecific with G. (s.str.) kashmirensis . Thus, I synonymyzed G. karakorus with this species.

Figures

FIGURE 51. Distribution of Geodromicus kashmirensis in the Central Asia.

FIGURES 40–45. Habitus of Geodromicus: 40—G. kashmirensis (lectotype), 41—G. similis (lectoype), 42—G. consors (holotype), 43—G. karakorus (holotype), 44–45—G. lebedevianus (Fig. 44: Kyrgyzstan, Chichkan Vallley; Fig. 45: Kyrgyzstan, Kara-Balta). Scale bars: 1.0 mm.

FIGURES 46–50. Aedeagus of Geodromicus kashmirensis (Fig. 46: Afghanistan, Pagman; Fig. 47: India, Gulmarg; Fig. 48: India, Gya-Ladak; Figs. 49–50: India, Tangmarg Pir Panjal): 46–49—parameral view, 50—lateral view. Scale bars: 0.1 mm.

FIGURES 19–24. Aedeagus of Geodromicus zwickianus (Fig. 19: Iran, Qotur; Fig. 20–21: holotype of G. iranicus; Fig. 22: Iran, Kandovan Pass; Fig. 23: Iran, Zagros Mts.; Fig. 24: Iran, Küh-e Hezâr): 19–20, 22–24—parameral view, 21—lateral view. Scale bars: 0.1 mm.