Geodromicus (s.str.) curvipes Cameron, 1924

Shavrin, Alexey V., 2022, The bodemeyeri and the convexicollis species groups of the genus Geodromicus Redtenbacher, 1857, and additional taxonomic and faunistic data for some species of the Eastern Palaearctic Region (Coleoptera: Staphylinidae: Omaliinae), Zootaxa 5213 (5), pp. 451-496 : 489-490

Geodromicus (s.str.) curvipes Cameron, 1924

( Figs. 70 , 74 , 80–81 )

Geodromicus curvipes Cameron, 1924: 173 ; Cameron 1930: 164

Type material examined. Lectotype (here designated) of Geodromicus curvipes Cameron, 1924 , ♂ ( Fig. 74 ): ‘Type | H.T.’ <round label with red margin, printed>, ‘Gahan 7000`| Simla Hills. | Dr. Cameron | IX. 1921.’ <printed>, ‘ Geodromicus | curvipes | Cam’ <handwritten>, ‘M.Cameron. | Bequest. | B.M. 1955-147.’ <printed>, ‘ LECTOTYPE | Geodromicus | curvipes Cameron, 1924 | Shavrin A.V. des. 2022’ <red, printed>, ‘ Geodromicus (s.str.) | curvipes Cameron, 1924 | Shavrin A. det. 2022’ <printed> ( BMNH). Paralectotype, ♂ (dissected): ‘Gahan 7000`| Simla Hills’ <printed>, ‘Dr. Cameron | IX. 1921.’ <printed>, ‘ Geodromicus | curvipes | Cam’ <handwritten>, ‘M.Cameron. | Bequest. | B.M. 1955-147.’ <printed>, ‘ Geodromicus (s.str.) | curvipes Cameron, 1924 | Shavrin A. det. 2022’ <printed> ( BMNH).

Redescription. Measurements (n=2): HW: 0.90–0.95; HL: 0.62; OL: 0.25; TL: 0.17–0.19; AL (lectotype): 2.50; PL: 1.11–1.12; PWmax: 1.12; PWmin: 0.78–0.85; ESL: 1.36–1.42; EW: 1.49–1.54; MTbL (lectotype): 0.90; MTrL (lectotype): 0.57 (MTrL 1–4: 0.30; MTrL 5: 0.27); AW: 1.48–1.55; AedL: 0.87; BL: 4.80–4.95 (lectotype).

Habitus as in Fig. 74 . Body, mouthparts, antennae and legs dark-brown; tarsi yellow-brown. Head with fine microsculpture: transverse in clypeus and isodiametric in middle portion; neck with very dense isodiametric meshes; pronotum with very fine isodiametric microreticulation, indistinct in mediobasal portion; scutellum with fine transverse microsculpture; abdomen with dense and transverse meshes. Pubescence of forebody very fine, semierect, moderately short and sparse, longer in apical portion of head; abdomen with moderately dense and very fine, decumbent pubescence.

Head 1.4–1.5 times as broad as long, with distinctly convex supra-antennal elevations, middle portion between anterior margin of eyes and infraorbital portions; anterio-median depression subrectangular, wide and moderately deep; temples very convex, 1.3–1.4 times as long as longitudinal length of eyes; interocellar depression narrow, subtriangular and moderately deep, with narrow and deep anteocellar foveae, reaching level of apical third of eyes. Eyes large, strongly protruded laterad. Ocelli large, distance between ocelli slightly shorter than distance between ocellus and posterior margin of eye. Punctation moderately large and deep, sparser and finer in middle portion. Maxillary palpomere 3 slightly broadened apically; apical palpomere slightly shorter than 3, from middle gradually rounded toward rounded apex. Antenna reaching middle of elytra when reclined; antennomere 2 about 1.3 times as long as basal antennomere, 3 distinctly longer than 2, 4 shorter than 3, 5–8 slightly longer and broader than 4, 9–10 slightly broader than 8, apical antennomere 1.3 times as long as 10.

Pronotum convex, about as broad as long, from widest middle strongly narrowed anteriad toward rounded apical angles and gradually narrowed toward base; narrowest basal part of pronotum slightly elongate, indistinctly concave laterally and with slightly divergent lateral subacute hind angles; middle portion with long and moderately wide longitudinal depression (lectotype with depression reaching middle length of pronotum; longitudinal depression of paralectotype reaching mediobasal portion of pronotum); mediobasal depression transverse and moderately deep; apical portion of pronotum slightly protruded apicad, with anterior margin widely concave, about as broad as slightly concave basal margin, with bordered and slightly reflexed edges, larger in latero-apical portions; lateral margins of pronotum narrowly bordered. Punctation about as that in middle portion of head, but slightly denser, significantly sparser and finer in mediobasal portion in front of mediobasal depression

Elytra convex, depressed in middle, slightly broader than long and slightly broadened posteriad, 1.2 times as long as pronotum; hind margins widely rounded. Punctation denser, coarser, larger and deeper than that on pronotum, with distance between punctures in middle as long as diameters of one–two nearest punctures, finer along suture (paralectotype) and/or apical portions.

Metatibia about 1.5 times as long as metatarsus.

Abdomen about as broad as elytra, with two large and transverse tomentose spots in middle of abdominal tergite IV and moderately wide palisade fringe on apical margin of abdominal tergite VII.

Male. Profemuri very wide; inner margins of protibia widely curved; protarsomeres 1–4 moderately wide. Apical margin of abdominal tergite VIII truncate or slightly concave. Apical margin of abdominal sternite VIII widely concave. Aedeagus with wide basal part, significantly narrowed toward median lobe; median lobe with elongate and moderately wide apical portion, from apical third gradually narrowed toward rounded apex; parameres narrow and short, not reaching apex of median lobe, with three moderately short apical and two subapical setae; internal sac with moderately short and wide field of small thorns, with elongate, sclerotized, apical structures and very long, narrow flagellum ( Fig. 80 ). Lateral aspect of the aedeagus as in Fig. 81 .

Female unknown.

Comparative notes. Based on the general shape and the coloration of the body with similar shape of the pronotum, the presence of microsculpture and longitudinal depression on the pronotum, the similar shape of the protibia in males and the general shape of the aedeagus, G. (s.str.) curvipes is similar to the Himalayan G. (s.str.) subsimilis (see below). From this species it can be distinguished by the smaller body, slightly narrower pronotum, finer microsculpture on the pronotum, broader apical portion of the median lobe and different internal morphology of the aedeagus, with the presence of the long flagellum (missing in G. (s.str.) subsimilis ).

Distribution. The species is known from the type localitiy in Himachal Pradesh, India ( Fig. 70 ).

Bionomics. Specimens were collected from 2100 m a.s.l. The detailed bionomical data are unknown.

Remarks. Geodromicus curvipes was originally described based on unspecified number of specimens from “Simla Hills; Gahan, 7000 feet above the sea”. The male in better condition and with additional label ‘Type H.T.’ is designated as the lectotype in order to fix the identity of the name.

Figures

FIGURE 70. Distribution of Geodromicus in the Himalayan Region: G. affinis (circles), G. alticola (squares), G. curvipes (triangle), G. ruficornis (diamond), G. subsimilis (black stars).

FIGURES 74–77. Habitus of Geodromicus: 74—G. curvipes (lectotype), 75—G. ruficornis (holotype), 76—G. subsimilis (holotype), 77—G. smetanai (China, Hailuogou). Scale bars: 1.0 mm.

FIGURES 78–83.Aedeagus of Geodromicus alticola (Figs. 78–79: India, Lianmarg), G. curvipes (Figs. 80–81: paralectotype), G. subsimilis (Figs. 82–83: India, Mangpo): 78, 80, 82—parameral view, 79, 81, 83— lateral view. Scale bars: 0.1 mm.