Hanakia HORÁČEK , 2001

Rosina, Valentina V. & Rummel, Michael, 2019, The Early Miocene Bats (Chiroptera, Mammalia) From The Karstic Sites Of Erkertshofen And Petersbuch 2 (Southern Germany), Fossil Imprint 75 (3 - 4), pp. 412-437 : 421

Genus Hanakia HORÁČEK, 2001

Hanakia agadjaniani ROSINA et RUMMEL, 2012 Figs 3a–g, i, l, 4

M a t e r i a l. Erkertshofen 2: BSP 1974 XIV 1194, left C sup.; BSP 1974 XIV 1195, right C sup.; BSP 1974 XIV 1192, right mxl with P4–M3; BSP 1974 XIV 1193, right mxl with P4–M2; BSP 1974 XIV 1197, right P4; BSP 1974 XIV 1198, right M1; BSP 1974 XIV 1196, right C sup.; BSP 1974 XIV 993, left mnd with talonid m2 and m3; BSP 1974 XIV 994, right mnd with m3; BSP 1974 XIV 995, right mnd with m2–3; BSP 1974 XIV 1189, right mnd with m2; BSP 1974 XIV 1191, left m1; BSP 1974 XIV 1190, left C inf.; BSP 1974 XIV 1111, right m1.

Petersbuch 2: BSP 1976 XXII 5512a–q, PCMRCh93a–j, PCMRCh94a–h, PCMRCh95a–i, PCMRCh96a–f, PCMRCh97a–f, PCMRCh98a–h, PCMRCh102a–c, PCMRCh114, PCMRCh8, PCMRCh9 (66 isolated upper teeth); BSP 1976 XXII 5488–5489, BSP 1976 XXII 5493–5497, PCMRCh90–92 (10 maxillary fragments with teeth); BSP 1976 XXII 5490–5492, PCMRCh10–11, BSP 1976 XXII 5498–5503, BSP 1976 XXII 5509h, BSP 1976 XXII 5511d–f, PCMRCh103a–i, PCMRCh104a–c (23 mandibles with teeth); BSP 1976 XXII 5508a–o, BSP 1976 XXII 5509a–g, BSP 1976 XXII 5510a–c, BSP 1976 XXII 5511a–c, PCMRCh99, PCMRCh100a–d, PCMRCh101, PCMRCh104d–f, PCMRCh105a–h, PCMRCh106a–g, PCMRCh107a–u (73 isolated lower teeth).

M e a s u r e m e n t s. See Tab. 5.

D e s c r i p t i o n a n d c o m p a r i s o n. These specimens from Erkertshofen 2 and Petersbuch 2 show all the typical features of the genus Hanakia , particularly the robust construction of the cranium while the dental formula corresponds to Myotis . However, in its dentition the talonid of m3 is considerably reduced and the incisive row is markedly compressed. The mandible is robust and the preserved alveoli of i1–3 suggest that the incisors were similar in size, positioned very close to each other and that the i3 was buccally displaced (e.g. BSP 1976 XXII 5490; Text-fig. 4b ). The ascending ramus is nearly at right angles to the body of the mandible and the masseteric fossa is very deep (e.g. BSP 1976 XXII 5492; Text-fig. 4d ). The lower canine has a notable mesolingual cingular cuspid and is surrounded by a well-developed cingulum (e.g. PCMRCh10; Text-fig. 4c ). The crown of the p2 has two small anterior and posterior tubercles. The p4 is quadrangular in occlusal view, surrounded by a distinct cingulum and has small anterior and posterior tubercles on the lingual side of the crown (e.g. BSP 1976 XXII 5491; Text-fig. 4e ). All lower molars are myotodont with distinctly curved paralophids. The talonid of the m3 is markedly reduced (e.g. PCMRCh11; Text-fig. 4f ). Hanakia from Erkertshofen 2 and Petersbuch 2 differs from H. fejfari from Merkur-North (= Ahníkov, MN 3, Czech Republic; Horáček 2001) in being smaller ( Tab. 6) and in having larger p2–3 (compare with Horáček 2001: 139, figs 18, 19). The morphology of Hanakia from Erkertshofen 2 and Petersbuch 2 is almost identical to that of H. cf. agadjaniani (“ Myotis cf. antiquus ”; for details see Rosina and Rummel 2012) from Wintershof-West ( Ziegler 1993) and from Stubersheim 3 ( Ziegler 1994). However, the fossils under study differ from H. cf. agadjaniani in having a less reduced premolar tooth-row but a more robust mandibular body, in having a more reduced m3 talonid and M3, and in a lack of the additional conules and lophs on the upper molars ( Text-figs 3 , 4 ). Hanakia from Erkertshofen 2 and Petersbuch 2 differs from the H. antiquus (La Grive 7, MN 7, France, Gaillard 1899) in having a less reduced premolar tooth-row and a more robust mandibular body (Hmdm1 of M. antiquus is only 1.5 mm; compare with Tab. 5). Moreover, the m3 talonid of H. antiquus is only slightly reduced in size ( Gaillard 1899, Viret 1951: 18, fig. 3) also distinguishing this species from the specimens under study. Its M1–2 have para- and metalophs ( Baudelot 1972: 50, tab. 2) and the M3 is less reduced ( Viret 1951: 18, fig. 2) in contrast to the fossils from Erkertshofen 2 and Petersbuch 2. Morphologically these fossils from Erkertshofen 2 and Petersbuch 2 correspond to the type specimens of H. agadjaniani from Petersbuch 62 ( Text-figs 3 , 4 ; Rosina and Rummel 2012: 468–470). There are remains of another form of Hanakia in the localities of Petersbuch 2, Erkertshofen 1 and Erkertshofen 2, which are noticeably smaller in size than H. agadjaniani from Petersbuch 2 and Erkertshofen 2 and differs in some other morphological features (see below).

Figures

Text-fig. 4. Lower jaw fragments. Hanakia agadjaniani: a – PCMRCh101, left p2, Petersbuch 2, occlusal view; b – BSP 1976 XXII 5490, left mnd with m2–3, Petersbuch 2, occlusal (b1) and lateral (b2) views; c – PCMRCh10, right mnd with C inf.–p2, Petersbuch 2, lingual (c1) and occlusal (c2) views; d – BSP 1976 XXII 5492, right mnd with m2, Petersbuch 2, lateral view; e – BSP1976 XXII 5491, right mnd with p4–m1; Petersbuch 2, occlusal view; f – PCMRCh11, left mnd with m1–3, Petersbuch 2, occlusal view.

Text-fig. 3. Upper jaw fragments. a–g, i, l – Hanakia agadjaniani: a – PCMRCh90, left mxl with M1–3, Petersbuch 2, ventral view; b – PCMRCh91, left mxl with M2–3, Petersbuch 2, ventral view; c – BSP 1976 XXII 5488, right mxl with M1–3, Petersbuch 2, ventral view; d – BSP 1974 XIV 1192, right mxl with P4–M3, Erkertshofen 2, ventral view; e – PCMRCh9, right C sup., Petersbuch 2, lingual view; f – PCMRCh8, right C sup., Petersbuch 2, occlusal view; g – PCMRCh99, left P2, Petersbuch 2, occlusal view; i – PCMRCh114, left M2, Petersbuch 2, occlusal view; l – BSP 1976 XXII 5489, left mxl with P2–M2, Petersbuch 2, ventral view; h – H. agadjaniani, NMA P62/0334, right mxl with P2–M2, Petersbuch 62, ventral view; j, k – H. aff. antiquus, Petersbuch 2: j – PCMRCh2, left M1, occlusal view; k – PCMRCh1, left M2, occlusal view.