Anthoxyela orientalis, Gao, Taiping & Ren, Dong, 2008

Gao, Taiping & Ren, Dong, 2008, Description of a new fossil Anthoxyela species (Hymenoptera, Xyelidae) from Yixian Formation of Northeast China, Zootaxa 1842, pp. 56-62 : 58-60

Anthoxyela orientalis sp. nov

( Figs. 1–3 )

Diagnosis: This species can be diagnosed as follows: Forewing with basal section of Rs approximately equal to M; hind wing with Rs approximately vertical to R1, and basal part of vein cu-a extraordinarily bend; pronotum in dorsal view shorter, posterior margin slightly concaves.

Description: Color dark, particularly on head; metascutum and abdomen dark brownish, antenna and legs slightly light brownish.

Head large and round, clypeus broad and margin round; mandible triangular with slightly elongate apex, intercrossing, eyes large, kidney-shaped, ocelli clear and display appreciably as an obtuse angle. Antennae, as preserved, about twice longer than the width of head, terminal part of flagellum at least thirteen segments, each with length slightly longer than width, the third segment thick and almost equal to rest flagellomeres, but thinner than the pedicel. Pronotum short, trapeziform. Mesonotum with median suture clear, anterior margin round. Abdomen with 8 segments visible; and genitalia partly absent, volsellaes digitiform, extending to the seventh abdominal segment.

Forewing with pterostigma sclerotized basally only, costal area obviously widen at the base of Rs, Sc meeting R only a very short distance beyond origin of Rs, posterior branch of Sc(Sc2) short and subvertical to R, first segment of Rs slightly longer than M; R curve at the origin of Rs, and greatly thickened before pterostigma; 1r-rs connecting to R just at the base of the break adjacent to pterostigma; 1r-rs and 2r-rs approximately equal in length; the segment of M between 1m-cu and 2r-m approximately straight; crossvein 1m-cu shorter than the segment of Cu toward it and slightly longer than 1cu-a; cell R 1.5 times as long as 1R1; cell 1M equal to 1R1, and 2.4 times longer than wide; cell 2M equal to cell Rs in length; cell 1Cu narrow and distinctly broad at the origin of vein M, 6 times longer than wide, and 2.5 times than cell 1M in length.

Right hind wing partly visible, preserved behind the forewing, the basal section of Rs longer and subvertical to R1, and basal part of vein cu-a extraordinarily bend, R1 thicker at basal part, hamuli visible, cell 1Cu broad.

Measurements (mm): Length of body without male genitalia 15.1mm, forewing 10.2mm as preserved, and antennae not shorter than 5.2mm, head 2.9mm in length and 2.8mm in width, thorax 3.8mm and abdomen preserved 10.0mm.

Holotype: No. CNU-HYM-LB-2008003, an impression of a nearly complete male sawfly in dorsal view. It is housed in the Key Lab of Insect Evolution and Environmental Changes, College of Life Science, Capital Normal University, Beijing, China.

Etymology: From the Latin orientalis referring to the locality ( China) of the fossil site.

Locality and horizon: Yixian Formation, Huangbanjigou, Chaomidian Village, Beipiao City, Liaoning Province, Northeast China.

Remark: The new species differs from A. baissensis, Rasnitsyn, 1977 , in shorter basal segment of the Rs, and shorter Sc2; differs from A. anthophaga Rasnitsyn, 1982 , in its shorter pronotum and length of 1m-cu against adjacent sections of M and Cu. In A. turgensis, Rasnitsyn, 1990 , the base of Rs is shorter than the first segment of M, and the anteriorly branch of Sc is slightly longer, these characters can distinguish A. turgensis, Rasnitsyn, 1990 from the new species.

Figures

FIGURE 1. Anthoxyela orientalis sp. nov., photograph of holotype, CNU-HYM-LB- 2008003, dorsal view.

FIGURE 2. Anthoxyela orientalis sp. nov., line drawing of holotype CNU-HYM-LB- 2008003.

FIGURE 3. Anthoxyela orientalis sp. nov., line drawing of holotype CNU-HYM-LB- 2008003. A. right forewing; B. head; C. antenna.