Acalanthis quadrisignata Erichson

Arias, Elizabeth T., Slipinski, Adam, Lawrence, John F. & Elgueta, Mario, 2009, A review of the Chilean Egoliini (Coleoptera: Trogossitidae) with description of a new species of Necrobiopsis Crowson, Zootaxa 2170, pp. 37-45 : 39-40

Acalanthis quadrisignata Erichson

( Fig. 3 )

Acalanthis quadrisignata Erichson, 1842: 446 .

Psoa quadrinotata Blanchard, 1851: 436 –437, 1854: pl. 26, fig. 7. Synonymized by Lacordaire, 1854: 336.

Diagnosis: This species is easily recognised by its relatively dull body with each elytron bearing 10 longitudinal rows of punctures forming distinctly impressed striae and two amber-colour spots; other diagnostic features include relatively small eyes, the pronotum widest at anterior angles, weakly and more or less evenly narrowing posteriorly, and the surfaces of head and pronotum entirely, coarsely punctate.

Description: Length 5.7–9.5 mm. Colour pitch black, legs and antennae somewhat lighter; each elytron with two amber colour transverse spots ( Fig. 3 ); vestiture consists of moderately long, erect, yellow setae of variable lengths with the longest setae situated mostly along lateral parts of head and pronotum, elytral intervals and lateral margins.

Head, including eyes, as wide as pronotum at anterior angles; punctation very coarse and dense, covering entire head surface; interspaces densely reticulate and dull. Terminal maxillary palpomere, three times as long as wide and two times as long as penultimate. Antennal club 1.3 times as long as wide and two times as long as penultimate segment.

Pronotum 1.1–1.2 times as long as wide, widest at anterior angles or shortly behind, and weakly narrowing posteriorly; lateral margin complete, crenulated, without distinct bead. Disc evenly convex, without impressions or tubercles, entirely covered by coarse and dense puctation with punctures about as large as those on vertex; interspaces densely reticulate, mat or feebly shiny. Scutellum pentagonal, transverse and distinctly elevated.

Elytra 2.2–2.5 times as long as wide and 2.5–2.6 times as long as pronotum; sides parallel for most of the length with both elytra jointly rounded apically; disc evenly convex and feebly shiny at most. Each elytron with 10 complete rows of strongly elongate, somewhat continuous punctures, giving appearance of impressed lines or grooves; intervals flat, slightly wider that “striae” and sparsely setose.

Specimens examined: Chile: Talinay Coquimbo, 21.iv.1955, L. Peña (1, ANIC); Coquimbo, Talinay, 16.x.57, s/ Drymis winteri, G. Kuschel (1, MNNC); Coquimbo, Talinay, 16.10.57, ex Rhaphithamnus spinosus, G. Kuschel (1, MNNC); Talinai, Coquimbo, 27.i.1969, J. Solervicens, en tronco olivillo (6, UMCE); Limarí, Talinay, 13.x.1976, J. Solervicens (1, UMCE); Limarí, Talinay, 25.x.89, J. Solervicens (7, UMCE); Limarí, Talinay, 26. x.1989, J. Solervicens (6, UMCE); Limarí, Talinay, 28.x.89, J. Solervicens (1, UMCE); Villa Alemana, Tranque Recreo, 18.ii.1974, J. Solervicens (1, UMCE); Laguna El Peral, 2. Jun. 1946, G. Kuschel, ex Schinus latifolius (1, MNNC); Talagante, Talagante, 6.viii.1992, J. Mondaca (2, MNNC); Santiago, Cerro Chena, 15.viii.1975, G. Arriagada (1, MNNC); Santiago, Cerro Cantillana, 15-16.xii.1979, J. Solervicens. 2100 m. En tronco roble (2, UMCE); Cardenal Caro, Quebrada El Roble, 28.i.89, J. Solervicens (1, UMCE); Curicó, Puente Quebrada Mala, 32 km interior Molina, 2.xi91, J. Solervicens (5, UMCE); Talca, Altos de Vilches, 1.280 m. 26.i.69. Collector J. Valencia (1, UMCE); Altos de Vilches, Talca 1.280 mts., 25- 26.xi.1970, J. Solervicens (1, UMCE); Talca, Altos de Vilches, 1.xi.1971, Col. C. Vivar T. (1, UMCE); Talca: Alto Vilches, 1100 m, 10-12.xii.1976, H.F. Howden (1, ANIC); Vilches Alto, x.1990 (1, ANIC); Chile Talca, Vilches, 10.ii.2007, Luis Flores (1, UMCE); Chile Linares, Estero Leiva, 8-12.i.1953, L. Peña (3, MNNC); VIII Region, N Los Lleuques/ Puente Marchant, 36°54.388’S / 071°32.111’W, 1022 m, 2.xii.2001, Canopy fogging GT, Nothofagus dombeyi, Arias et al. UC Berkeley (7, EMEC); Concepción, Florida km. 40, 6.xii.89, J. Solervicens (1, UMCE); Lanalhue, Arauco, 10.x.1969, J. Solervicens (4, UMCE); Malleco: 20 km E Manzanar, 1100 m, 19-21.xii.1976, H.F. Howden (1, ANIC); IX Región Prov. Malleco, Malalcahuello, Feb. 2004, R. Barrera (1, MNNC); IX Region, Nahuelbuta NP exit, Pichinahuel Fogging, 05.i.2007, 1238 m 22o C, A. araucana 15 m, S37 o49.041/ W73 o0 0.214, Arias et al Berkeley (4, EMEC); Cherquenco, i-ii.1954, T. Ramirez (1, ANIC); Valdivia, Puerto Pirihueico, 6.ii.90, J. Solervicens (1, UMCE); Provincia Valdivia, Valdivia, 21.x.1971, E. Krahmer (1, MNNC); same, 28.x.1971, E. Krahmer (1, MNNC); Valdivia: 30.i.1975, L. Peña (1, ANIC); Puerto Varas, i.1924 (1, MNNC); Petrohue, Llanquihue, 24.ii.1969, J. Solervicens (1, UMCE); Chiloe, Archipielago Guapiquilan, Isla Leguas, 26.i.87, J. Solervicens (1, UMCE); Chiloe, Isla Quilán, 21- 25.i.87, J. Solervicens (1, UMCE); Osorno, Lago Puehue, 2 km W Termas de Puyehue, 40?42'S.72?22W, 300 m, 13.xi.1994, R. Leschen, C. Carlton, #091, ex beating vegetation (R. Leschen, collection).

Distribution: Chile. Some specimens have been collected in bog forests with several species of Myrtaceae and Drimys winteri J.R. Forst. & G. Forst. (Winteraceae) ( Solervicens & Elgueta, 1994).

Notes. Although the synonymy of Psoa quadrinotata with Acalanthis quadrisignata is a well established one, which has been recognized in several later works ( Reed, 1876: 277; Philippi, 1887: 673; Léveillé, 1889: 431; Léveillé, 1910: 4; Blackwelder, 1945: 391), the name has been recently cited in error as a valid species of Bostrichidae by Borowski & Węgrzynowicz (2007).

Figures

FIGURES 1 – 4. Adult habitus, dorsal. 1. Acalanthis semimetallica (Fairmaire & Germain); 2. A. mirabilis Reitter; 3. A. quadrisignata Erichson; 4. Necrobiopsis shangrila sp. nov.