Marmessoidea alata, Ho, 2018

Ho, Wai-Chun George, 2018, New taxa of Necrosciinae from Vietnam (Phasmatodea: Diapheromeridae), Zoological Systematics 43 (2), pp. 178-189 : 184-185

Marmessoidea alata alata sp. nov. & subsp. nov. ( Figs 13–16 , 30–31 )

Diagnosis. This new species is related to M. haemorrhoidalis Redtenbacher, 1908 , but can be diagnosed by green mesosternum and longer alae. The male is also related to M. liuxingyuei sp. nov., but can be separated by greenish body and uniformly coloured legs.

Description. Male. Body slender, distinctly smaller than female. General colour of body and legs dull green. Head oval, almost as long as pronotum. Vertex flat, with a few small granules. Occiput convex, with distinct median longitudinal furrow, posterior margin with indistinct swellings. Compound eyes rounded, about 2 times as long as genae. Antennae very long and filiform, longer than body length, covered with dense short bristles; scapus flattened, about 2 times as long as pedicellus; third segment longer than pedicellus, as long as scapus. Pronotum with sparse granulations, rectangular, longer than wide, anterior margin curved inwards, posterior margins rounded, longitudinal and transverse sulci crossing before middle area. Mesonotum with sparse granulations, almost parallel-sided, slightly constricted post-medially, almost as long as combined length of metanotum and median segment, with faint median longitudinal line. Mesopleurae, metapleurae, mesosternum and metasternum lacking granulations. Abdomen lacking granulations. Eighth tergum longer than ninth tergum. Anal segment as long as eighth tergum, with a small emargination on posterior margin. Poculum tapering posteriorly, posterior margin pointed. Cerci cylindrical, apices obtuse and gently curved inwards. Legs unarmed, slender and long, with short bristles. All femora roughly as long as corresponding tibiae. Profemora incurved basally. Tegmina oval, posterior margin subtruncate, with a black and blunt elevated angle near pre-medial area, also with a yellowish green marking near lateromedial area. Alae green, long and well-developed, posterior apices reaching sixth abdominal tergum.

Female. Body cylindrical, distinctly more robust and larger than male. Body and legs brown with light brown markings. Head oval, as long as pronotum. Vertex flat, with a few of minute granules. Occiput convex, with distinct median and lateral longitudinal furrows, posterior margin with indistinct swellings. Compound eyes rounded. Antennae long and filiform; scapus more or less flattened, longer than pedicellus, roughly as long as third segment. Pronotum with small granules, rectangular, longer than wide, with longitudinal and transverse sulci crossing before middle area, anterior margin curved inwards, posterior margin rounded. Mesonotum with small granules, moderately constricted post-medially, with distinct median longitudinal line. Mesopleurae, metapleurae, mesosternum and metasternum with small and inconspicuous granules. Abdomen with inconspicuous granulations, cylindrical, tapering posteriorly. Seventh sternum lacking praeopercular organ. Ninth tergum shorter than eighth tergum. Anal segment as long as eighth tergum, posterior margin with a small emargination. Supra-anal plate small, median longitudinal carina distinct, posterior margin rounded. Subgenital plate scoop-shaped, posterior margin with a deep U-shaped emargination. Cerci long, straight and cylindrical, apices pointed and surpassing posterior margin of anal segment. Legs unarmed, slender and long. Profemora incurved basally. Tegmina oval, longer than head. Alae short, posterior apices reaching anterior area of fifth abdominal tergum, anal region brown.

Measurements. Length. Body, ♀ 66.0–73.0 mm, ♂ 53.0–65.0 mm; antennae, ♀ 60.0–68.0 mm, ♂ 72.0–87.0 mm; head, ♀ 5.0 mm, ♂ 3.0–4.0mm; pronotum, ♀ 5.0 mm, ♂ 3.0–4.0mm; mesonotum, ♀ 14.0–15.0 mm, ♂ 10.5–11.5mm; metanotum (including median segment), ♀ 8.5–9.0 mm, ♂ 7.5–10.0 mm; profemora, ♀ 17.0–19.0 mm, ♂ 18.0–20.0 mm; mesofemora, ♀ 11.0–13.0mm, ♂ 11.0–14.0 mm; metafemora, ♀ 17.0mm, ♂ 17.0–21.0mm; protibiae, ♀ 16.0–19.0 mm, ♂ 16.0–20.0mm; mesotibiae, ♀ 10.0–13.0 mm, ♂ 11.0–15.0 mm; metatibiae, ♀ 16.0–18.0 mm, ♂ 16.0–22.0 mm; tegmina, ♀ 6.0– 7.5 mm, ♂ 4.0–6.0 mm; alae, ♀ 24.0–28.0 mm, ♂ 26.0–36.0 mm.

Material examined. Holotype ♂, Vietnam, Thua Thien Hue, Phu Loc, Bach Ma (elev. 150 m), 27 September 2011, Liu Xingyue ( CAU) . Paratypes. 1♂, 1♀ (subadult), Vietnam, Thua Thien Hue, Phu Loc, Bach Ma (elev. 150 m), 27 September 2011, Liu Xingyue ( CAU & HKES) ; 1♂, Vietnam, Thua Thien Hue, Bach Ma , 4 May 2012, Liu Xingyue ( CAU) ; 1♂, 2♀, Vietnam, Gia Lai, c. 40km N of K’ Bang Town, Kon Chu Rang Nature Reserve (14°30'19''N, 108°32'28''E; elev. 1020 m), 24 May–2 June 2016, Abramov A.V., F3476.23, F3476.28, F3476.39 ( MMUE & HKES) .

Distribution. Vietnam.

Etymology. The specific epithet of this new species is derived from the well-developed wings.

Figures

Figures 1–23. Necrosciinae spp. 1–4. Asceles hirsutus sp. nov. 5–8. A. maculatus sp. nov. 9–12. Lamachodes brocki sp. nov. 13– 16. Marmessoidea alata alata sp. nov. & subsp. nov. 17–18. M. liuxingyuei sp. nov. 19–23. Neooxyartes zomproi gen. nov. & sp. nov. 1, 5, 9, 15, 19. Female end of abdomen, lateral view. 2, 6, 10, 16, 20. Female end of abdomen, dorsal view. 3, 7, 11, 13, 17, 21. Male end of abdomen, lateral view. 4, 8, 12, 14, 18, 22. Male end of abdomen, dorsal view. 23. Vomer, ventral view. Scale bars: 1–22=5mm; 23=1mm.

Figures 30–35. Necrosciinae spp., habitus. 30–31. Marmessoidea alata alata sp. nov. & subsp. nov. 32. M. alata elongata subsp. nov. 33. M. liuxingyuei sp. nov. 34–35. Neooxyartes zomproi gen. nov. & sp. nov. 30, 32–33, 35. Male. 31, 34. Female. Scale bars= 5mm.