Panscopus (Nocheles) aequalis (Horn, 1876)

Spanton, Timothy G. & Anderson, Robert S., 2016, A Taxonomic Revision of Weevils of the Genus Panscopus Schönherr (Coleoptera: Curculionidae: Entiminae: Tropiphorini), The Coleopterists Bulletin (mo 15) 70, pp. 1-86 : 55-58

Panscopus (Nocheles) aequalis (Horn, 1876)

( Plate 6 , Figs. 23 , 49 , Map 24 )

Nocheles aequalis Horn 1876: 55 .

Nocheles cinereus Horn 1876: 55 [in key, lapsus calami for N. aequalis ].

Nocheles vestitus Casey 1888: 251 .

Panscopus (Neopanscopus) aequalis ; Pierce 1913: 397; Buchanan 1927: 28, 1936: 13; Schenkling and Marshall 1931: 40; Blackwelder 1939: 66; Bousquet et al. 2013: 345.

Panscopus aequalis ; Hatch 1971: 290; O’ Brien and Wibmer 1982: 62; Bright and Bouchard 2008: 254.

Type Material. Lectotype (MCZ No. 2818): “Col.”, indicating that it was collected in Colorado, and “Horn Coil. H 8279” “ N. aequalis H.”.

Specific Epithet. The Latin adjective “ aequalis ” means like, same, uniform. This could refer to the topographically even elytral intervals, or the almost uniform rows of fine setae on the elytra, which are characteristic features of this species.

Diagnosis. The combination of having all elytral intervals equally convex, the alternate intervals not raised above the even intervals, the thin, semierect, acute setae in rows on all elytral intervals, and the coarse rugose-tuberculate surface of the pronotum, with tubercles visible through the vestiture, distinguish specimens of P. aequalis from all other species of Panscopus with the exception of P. longus . The latter and P. aequalis can be separated by the characters given in the key.

Redescription. Females: Length 6.6–8.7 mm (n = 10); width across elytra 2.6–3.5 mm (n = 10). Males of this species are unknown. Color: Grey to grey-brown. Head: Transverse impression at base of frons slight; frons and rostrum apparently nearly continuous in lateral view; vestiture of frons and rostrum of dense, overlapping grey scales and decumbent, stout, blunt setae; nasal plate indistinct; nasal carina obvious, v-shaped. Antennae: Scape short, extending to only anterior 1/4 of eye, with dense vestiture of scales and coarse setae; integument obscured over most of length. Pronotum: Approximately 1/10 wider than long, lateral margins gently rounded; median longitudinal sulcus incomplete and indistinct, more obvious in anterior half; dorsal surface coarsely rugose; vestiture of overlapping grey scales and scattered, decumbent setae. Elytra: broadly oval in dorsal view, combined width approximately 1/2 wider than pronotum, and approximately 72% of elytra length; declivity in lateral view, broadly, continuously rounded to reflexed apex; all elytral intervals equally slightly convex, each interval with complete row or partially double row of semi-erect, fine, acute setae; serial punctures each with scale, scales of serial punctures oval, in some slightly angular apically; elsewhere vestiture of pale grey scales or with patches of beige, overlapping, striate scales. Legs: Tibiae 1, 2, and 3 with row of stout spines on ventral edge in distal 1/3, more pronounced on tibiae 1 and 2; tibia 3 slightly bent, corbellar area closed. Abdomen: Ventrite 1 of females slightly convex at middle; ventrites with vestiture of grey scales similar to those of dorsum. Genitalia: Females ( Figs 23 , 49 ): Sternum VIII (spiculum ventrale) with apical expansion approximately 1/3 its length; coxites angular in dorsal view; vagina membranous, with pair of crescent-shaped sclerites near confluence with common oviduct; spermatheca sickle-shaped, broad, tapered toward apex in distal 1/2.

Life History. Of the 47 specimens examined that bear collection dates on labels, most were collected in May and June, a lesser number in July and August, and a few in March and April.

Plant Associations. Plant associations taken from label data are as follows: 19 specimens from Artemisia rothrockii A. Gray , four specimens from rabbitbush ( Chrysothamnus sp. , Asteraceae ), four specimens from rose ( Rosa sp. ), three from Salix sp. , and one from each of P. tremuloides and alfalfa ( Medicago sativa L., Fabaceae ).

Geographical Distribution. This species, of the Basin and Range region of interior western North America, exhibits the broadest range, both geographically and altitudinally, of any species of Panscopus . The range of P. aequalis is bounded in the west by populations in eastern and northern California, eastern Oregon and Washington, and the dry valleys of southern British Columbia, extending east across southern Alberta, Saskatchewan, and southwestern Manitoba, to North Dakota, Montana, Idaho, Wyoming, Kansas (doubtfully), Colorado, Utah, and Nevada ( Map 24 ). Altitudinal records extend from 910 m near Strathmore in southern Alberta to 3,200 m on the arid east side of the Sierra Nevada of California. Altitudinal records from localities in Idaho, Montana, Wyoming, and

Oregon vary from 1,520 to 2,290 m. An isolated record is from 2,900 m in the Ruby Mountains of northeastern Nevada.

Chorological Affinities. The range of this species overlaps completely the range of P. squamifrons .

Material Examined. In addition to the type material listed above, approximately 525 specimens were examined. CANADA: Alberta: Calgary , 3.V.1953, B.F. & J.L. Carr (1, CNCI) ; Calgary , 16.VI.1956, B.F. & J.L. Carr (1, CNCI) ; Calgary , Can. N.W.T., 17.V., ex Wickham collection. (1, USNM) ; Cardston, H.L. Seamans (1, CNCI) ; Cypress Hills , 10.VI.1932, O. Bryant (1, CASC) ; Cypress Hills , 2. VI.1956, B.F. & J.L. Carr (1, CNCI) ; Drumheller , 21.VI.1954, B.F. & J.L. Carr (1, CNCI) ; Edmonton , 26.VI.1919, F.S.Carr (1, CNCI) ; Empress , 11.2 km S. AB / SK border, 14.V.1973, 20.V.1973, B.F. & J.L. Carr (2, CNCI) ; Ghost Dam , 5.VI.1973, 3.VII.1974, 26.VI.1978, 15.V.1985, B.F. & J.L. Carr (4, CNCI) ; Gleichen , 8.V.1955, B.F. & J.L. Carr (2, CNCI) ; Highwood River , 31.V.1953, B. F. & J.L. Carr (1, CNCI) ; Hilda , 7.V.1930, A.T. McClay collection (3, UCDC) ; Lacombe , 9.VI.1924, R.D. Bird (1, CNCI) ; Lethbridge , 23.V.1930, 12.VII.1930, 19.VI.1930, 3.VII.1929, J.H. Pepper (4, CNCI) ; Canadian Forces Base Suffield , 50°37.678N, 110°18.371W, 16.VII.1994, Pantrap site 2.1.1, A.T. Finnamore (1, CMNC) ; Medicine Hat , 1 ( CWOB) ; Medicine Hat , 4.IV.1931, 14.VI.1931, A.T. McClay (3, UCDC) ; Medicine Hat , 14.VII.1925, 2.VI.1927, 21.IV.1930, 17.V.1930, 25.V.1930, 25.VI.1930, A.C. Davis (16, CNCI) ; Medicine Hat , 9.V.1926, Andrews (3, KSUC) ; Medicine Hat , 10.VI.1973, 5.VIII.1979, B.F. & J.L. Carr (2, CNCI) ; Medicine Hat, various dates-V-VII, 1921-1927, F.S. Carr (1, AMNH; 7, CNCI; 1, CWOB; 1, DEUN; 1, MTEC; 1, UCDC; 8, UAIC; 1, UMAA; 6, OSAC; 7, USNM; 2, CASC), Medicine Hat , 4-11.VII & 20.VII.- 4.VIII. 1980, G.A.P. Gibson, pan trap (5, CMNC) ; Medicine Hat , 15.VI.1930, J.H. Pepper (3, CNCI) ; Medicine Hat , 4.VI.1932, O. Bryant (4, CASC) ; Medicine Hat , 28.VI, O.H. Schwab collection (2, UCDC) ; Medicine Hat , 12-13.VI.1982, R.S. Anderson, pan trap (4, CMNC) ; Medicine Hat , 21.VI.1923, Shoemaker collection (10, USNM) ; Medicine Hat , 1.V.1923 (3, UMAA) ; Milk River , 19.V.1946, G.A. Hobbs (1, CWOB) ; Orion , 6.VI.1955, J.R. Vockeroth (2, CNCI) ; Orion , 3. VI.1922, H.E. Gray (1, CNCI) ; Patricia , 4.VIII.1923, H.L. Seamans (1, CNCI) ; Strathmore , 3.2 km W, 950 m, 16.VI.1962, C.W. O’ Brien (2, CWOB) ; Tilley , 10.VI.1932, O. Bryant (2, CASC) ; Tp. 1 Rge. 5, W 4 Mer. , 3.VII.1973, B.F. & J.L. Carr (1, CNCI) ; Tp. 2 Rge. 5, W. 4 Mer. , 18.V.1980, B.F. & J.L. Carr (1, CNCI) ; Tp. 8 Rge. 18, W. 4 Mer. , 21.V.1977, B.F. & J.L. Carr (1, CNCI) ; Tp. 11, Rge. 1, W. 5 Mer. , 8.V.1976, B.F. & J.L. Carr (1, CNCI) ; Tp. 13, Rge. 13, W. 4 Mer. , 24.V.1981, B.F. & J.L. Carr (1, CNCI) ; Tp. 15, Rge. 14, W. 4 Mer. , 1 0. V.1981, B.F. & J.L. Carr (1, CNCI) ; Tp. 21 Rge. 4, W. 4 Mer. , 30.VI.1973, B.F. & J.L. Carr (1, CNCI) ; Tp. 22, Rge. 1, W. 4 Mer. , 20.V.1973, B.F. & J.L. Carr (2, CNCI) ; Waiparous , 11.VII.1958, B. F. & J.L. Carr (2, CNCI) ; Sylvan Lake , 21.VI.1943, 28.VI.1943, E.J. Kitely (23, CNCI) . British Columbia: Aspen Grove , 13.V.1934, H.B. Leech, Populus tremuloides leaves (1, CASC) ; Aspen Grove , 19.V.1931, J.R. Howell (1, CASC) ; Aspen Grove , 14.VII.1922, 24.VI.1922, 29.VI.1922, P.N. Vroom (5, CNCI) ; Cawston , 3.VI.1917, 5.V.1917, 14.VI.1917, W.R.S. Metcalfe (3, CNCI) ; Chilcotin , 9.VI.1929, 14.VI.1931, G.J. Spencer (2, SMDV) ; Creston , 22.V.1959, G. Stace-Smith (1, SMDV) ; Kamloops , 1.V.1932, O. Bryant (5, CASC) ; Kamloops, Lac duBois, 18-22.VI.1954, G.J. Spencer (2, SMDV) ; Keremeos , 21.VI.1923, C.B. Garrett (1, CNCI) ; Marysville , 3.VI.1958, F. Fodor (2, SMDV) ; McLeod Meadows , 22.VII.1971, J.M. & B.A Campbell (1, CNCI) ; Merritt, Midday Valley , 29.V.1924, Culler (1, CNCI) ; Merritt, Midday Valley , 9.VI.1924, 15.VI.1924, 3.VII.1924, 15.VIII.1924, K.F. Auden (1, CNCI; 1, SMDV; 3, CASC) ; Merritt, Midday Valley , 6.V.1923, R. Hopping (1, CASC) ; Osoyoos , 22.V.1924, R. Hopping (1, CASC) ; Osoyoos, Richter Pass ”, 24.V.1959, L.A. Kelton (1, CNCI) ; Osoyoos, Richter Pass , 22.V.1959, R.E. Leech (2, CNCI) ; Penticton , 14.VI.1955, R. Stace-Smith (1, SMDV) ;. Princeton , 19.3 km E., 30.III.1941, H.B. Leech (1, SMDV) ; Princeton , 13 km E., 24.VIII.1977, K. Walde, steppe with Pinus ponderosa (1, UASM) ; Quesnel , 14.VII.1948, G.J. Spencer (1, SMDV) ; Radium , 4& 7.VI.1953, B.F. & J.L. Carr (2, CNCI) ; Spences Bridge , 18.V.1932, G. Stace-Smith (1, SMDV) ; Summerland , 20.IV.1942, S. Wilson (1, BMNH) ; Summerland , 3.VI.1932, 22.V.1931, A.N. Cartrell (2, CNCI) Summerland , 10.V.1906, J.R. Anderson (4, CNCI) ; Summerland , 20.IV.1942 (1, FMNH) . Manitoba: Aweme , 24.VI.1915, N. Criddle (1, CNCI) ; Treesbank , 19.V.1925, N. Criddle (1, CNCI) . Saskatchewan: Attons Lake, Cut Knife , VI.1940, A.R. Brooks (2, CNCI) ; Cypress Hills , 2.VI.1939, A.R. Brooks (3, CNCI) ; Earl Grey , 28. IV.1925, J. Ritchie (1, CNCI) ; Hague , 17.V.1924, K.M. King (1, CNCI) ; Oxbow , 16.V.1907, F.K. Knab (1, USNM) ; Rosthern , 16.V.1924, K.M. King (1, CNCI) ; Rutland , 2.VIII.1940, A.R. Brooks (1, CNCI) ; Saskatoon , 9.VI.1923, N.J. Atkinson (1, CNCI) ; Saskatoon , 26.IV.1927, 22.VI.1928, K.M. King (2, CNCI) ; Saskatoon , 1.VI.1906, T.N. Willing (1, CNCI) ; Val Marie , 13.VI.1955, J.R. Vockeroth (1, CNCI) . USA: California: Alpine Co., Markleeville , 5 km S.E., 1680 m, 10.VI.1966, C.W. O’ Brien (1, CMNC; 1, CWOB) ; Alpine Co., VII.1934, J.E. Blum (1, CASC) . Inyo Co., Crooked Creek Naval Research Station , 1.6 km W. White Mountains, VIII.1966, L. & C.W. O’ Brien, Artemesia rothrocki , night (17, CWOB) ; Crooked Creek Naval Research Station , 1.6 km W. White Mountains, 3200 m 4.VIII.1966, L. & C.W. O’ Brien, at night (3, CWOB) ; Crooked Creek Naval Research Station , 8 km S. White Mountains, 3050 m, 7.VII.1966, L. & C.W. O’ Brien (2, CWOB) ; Crooked Creek Naval Research Station , 8 km S. White Mountains, 3090 m, 20.VI.1961, G.W. Frankie (2, CWOB) ; Crooked Creek Naval Research Station , 1.6 km W. White Mountains, 3050 m, 4.VIII.1966, L. & C.W.O’ Brien (4, CWOB) ; Crooked Creek Naval Research Station, White Mountains , 3050 m, 4.VIII.1966, C.W. O’ Brien (18, CWOB) ; Vicinity of Rovana , 10.V.1999, J. Huether (1, CMNC) . Lassen Co.: Doyle , 31.V.1946, A T. McClay (1, UCDC) ; Hallelujah Junction , 28.V.1966, C.R. Kovacic (1, UCDC) ; Hallelujah Junction , 7.VI.1968, S.M. Hogue, R.L. Penrose, Salix (3, UICM) . Mono Co.: Bodie, Wickham (3, USNM) ; Bodie 2580 m, 1. VII.1907, Wickham (4, USNM) ; Bridgeport , 12 km W. 2100m, 6.VII.1966, C.W. O’ Brien (2, CWOB) ; Mono Lake , V.1920, 10.IV.1921, C. Hilton (2, CASC) ; Mono Lake 1940 m, 2.VII, Wickham (1, USNM) ; Sherman Creek , 29.VI.1955 (1, CWOB) ; Tom’ s Place 6.4 km S.W., 13.VII.1961, J.S. Buckett (1, UCDC) ; White Mountains . 3050 m, 23.VI.1961, D.R. Miller (1, UCDC) ; White Mountains, Blanco’ s Corral , 3050 m, 23.VI.1953, W.D. McLellan (2, UCDC) ; Mono Lake, Tioga Lodge , 22.VI.1929, E.P. Van Duzee (1, CASC) . Nevada Co., Boca , 3.VII.1954, R.C. Baylock (1, UCDC) ; Nevada City , 27.V.1941, A.T. McClay (2, UCDC) . Placer Co., Truckee 1770 m, Aug, Wickham (1, USNM) . Plumas Co., Graeagle , 15.VI.1949, E.L. Schlinger (1, UCDC) . Tuolumne Co., Chipmunk Flat , 9. VIII.1960, T. Gantenbein (2, UCDC) . County unknown: CA (1, USNM) : “ Cal. ”, H.W. Wenzel collection (2, OSUC) Rifle , 5.V.1905 (1, INHS) ; Tallac , 3.VII.1948, A. Fenyes collection (1, CASC) . Colorado: Jackson Co., North Sandhills Recreation Area , 29.V.2007, J. & B. Huether (1, CMNC) . Rio Blanco Co., Greasewood-Sage Site , #83A PF88 1950 m, VII.75, B.E. King, Rabbitbush (2, CWOB) ; Greasewood-Sage site #83A 1950 m, Aug-75, B.E. King, on Rabbitbush (2, CWOB) . Idaho: Bannock Co., 20.VI.1975, A.D. Allen (1, CWOB) . Blaine Co., Shoshone, 32 km N, Big Wood River , 1. VII.1966, W. Gagne, at night (1, CWOB) . Fremont Co., Island Park , 1.VII.1978, G.A. Shook (1, CIDA) . Nez Perce Co., Lewiston 1670 m, 15.IV.1936, V.E. Nygren (1, UICM) . County unknown: Id. , Horn (2, MCZC) ; Dillon Mountain. Idaho. Experiment. Station , 11.VII.1928, C. Wakeland (1, UICM) . Kansas: “ Ks ” (1, INHS) . Montana: Cascade Co., Cascade , 4.VIII.1952, taken from furniture (1, USNM) ; Cascade (1, USNM) . Gallatin Co., 1520 m, A.D. Hastings (1, USNM) ; Bozeman , 22.VI.1920, T.R. Chamberlain, Medicago sativa , swept (1, OSAC) ; Bozeman , 1.V.1927, 16.VI.1908 (2, MTEC) ; Gallatin Co., 8.VI.1954, Student collection (1, MTEC) . Granite Co., Phillipsburg , 17.7 km. S., 11.VII.1955, R.C. Froeschner (1, MTEC) . Ravalli Co., Hamilton , 18.VI.1928 (1, MTEC) . Roosevelt Co., Poplar , 19.VII.1956, R.C. Froeschner (1, MTEC) . Rosebud Co., Forsyth , 26.VI.1903 (1, USNM) . Yellowstone Co., Huntley , 11.VII.1914 (1, MTEC) . Counties unknown: “Mon”, Nason collection (2, INHS) ; “ Mont. ”, G. M. Gree. “ nt.”, Boulter collection (1, INHS) ; “ Montana ” (1, INHS) , 22.VI. (1, MTEC). Nevada: Elko Co., Carlin (1, USNM) ; Elko, Wickham (1, USNM) ; Ruby Mountains, Fauve Lake , 7,8 & 12.VII.1934, H.B. Stafford (4, EMUS) ; Fauve Lake, Ruby Mountains , 2900 m, 7& 12.VII.1934, H.B. Stafford (3, USNM) . Lander Co., Kingston , 6.VII.1993, Lot 4, B.F. and J.L. Carr (3, CMNC) . Washoe Co.: Verdi , 9.VII.1967, W. Gagne (4, CWOB) . Counties unknown: “Nev.”, Hubbard & Schwarz (2, USNM) ; “ Nev. ” (1, USNM) ; NV, 1.VII.1929, O.H. Schwab collection (1, UCDC) ; Carson City , 25.VI.1929, R.L Usinger (1, CASC) . North Dakota: Billings Co. Kadr. Dam, 10.VI.1945, D. Aarhus (1, NDSU) . Bottineau Co., Willow City , 17.VI.1977, R. Gordon (2, USNM) . McKenzie Co., Theodore Roosevelt. Memorial Park , 7.VI.1965, D. Aarhus, windowpane trap (1, NDSU) . Morton Co., Flasher Loc. No. 13, 13.VII.1966, R.J. Sauer, vacuum sweeper (1, NDSU) ; Wells Co.: III. (1, NDSU) . Williams Co.: Williston , 8-9.V.1919, Wickham (1, USNM) ; Slope 6, 13.VII.1965 (1, NDSU) . Oregon: Crook Co., Prineville , 30.VI.1946, A.T. McClay (1, UCDC) . Deschutes Co., Tumalo State Park , 22.VI.1954, K.M. Fender (1, OSAC) . Harney Co., Burns , 19.VI., B.G. Thomson (1, OSAC) ; Burns , 24 km. N. Hwy 395, 18.VI.1984, B.F. & J.L. Carr (1, CNCI) ; Fish Lake, Steens Mountains , 2160 m, 15.VII.1962 (1, USNM) ; Snake River, aids Ferry , 20.IV.1937, M.C. Lane (1, OSAC) ; Steens Mountains, Fish Lake , 2290 m, 20.VII.1955, J.E. Baker (1, AMNH) ; Steens Mountains, Fish Lake , 2290 m, 22-26. VI.1951, B. Malkin (3, OSAC) ; Steens Mountains, Fish Lake . 2290 m, 3& 4.VII.1947, F. Ellertson (2, OSAC) ; Steens Mountains, 6.4 km. W. Fish Lake , 15.VII.1953, Roth & Beer (1, OSAC) ; Steens Mountains, Fish Lake , 15.VII.1953, Roth & Beer (2, OSAC) ; Steens Goose Lake , 1923 (1, USNM) . Malheur Co., Harper , 21.V.1935, 10.V.1938, M.C. Lane (2, OSAC) ; Harper , 10.V.1938, E.W. Jones (3, UICM) ; Ironsides , 21.V.1935, M.C. Lane (1, OSAC) . Wasco Co., The Dalles , 11.3 km E., 10.IV.1969, R.L. Westcott, sandy habitat (2, ODAC) ; The Dalles , 21.III.1954, S. & M. Sargent (3, FMNH) . Utah: Beaver Co., Beaver Valley , 13. VI. (2, USNM) . Uintah Co., Vernal , 1.V.1952, N.R. Merrell (1, CWOB) . County unknown: “Ut”, J.B. Smith collection (2, USNM) Washington: Chelan Co., Chelan Falls, 26.IV.1911, Wash. Experiment Station #18 (1, USNM) ; Chelan Falls , 26.IV.1911 (1, WSU) . Grant Co., Grand Coulee , 2.IV.1933, J.F.G. Clarke (1, OSAC) ; Grand Coulee (1, OSAC) ; Grand Coulee , 2.IV.1933, I.W. Bates (4, WSU) . Okanagan Co., Brewster , 1 & 12.V.1912, Wash. Expt. Sta. #18 (2, USNM) ; Brewster , 1.V.1912 (1, WSU) ; Brewster , 17.VI.1954, D. Boddy & B. Malkin (2, FMNH) ; Okanagan , 4, 8, & 14. V.1912, Wash. Expt. Sta. #18 (1, USNM; 3, WSU) ; Okanagan , 8.V.1912 (1, BMNH) ; Omak , 3.V.1912 (1, USNM) ; Oroville , 10.V.1912 (1, USNM) ; Oroville , 24.VI.1912, Wash. Expt. Sta. #18 (2, WSU) ; Tonasket , 2.V.1911 (1, OSAC) ; Tonasket, M.A. Yothers (1, OSAC) ; Tonasket , 2.V.1911, Wash.Expt. Sta. #18 (2, USNM;18, WSU) ; Tonasket , 12.V. (22, WSU) ; Tonasket , 12. V., M.A. Yothers (7, WSU) . Walla Walla Co., Wallula , 28.V.1924, M.C. Lane (1, USNM) . Whitman Co., Hooper , 14.VI.1932, M.H. Hatch (1, OSAC) ; Pullman, C.V. Piper (1, USNM) ; Wawawai , 14.III.1910 (1, UCDC) ; Wawawai, C.V. Piper (7, USNM; 10, WSU) ; Wawawai , 8.V.1957, G. N. Knopf (1, UICM) . Yakima Co., Toppenish , 26.IV.1926, F.H. Shirck (1, OSAC) . Counties unknown: “Was.”, 26. IV.1911, Van Dyke collection (1, CASC) ; Mission , 5.V.1911, Wash. Expt. Sta. #18 (3, USNM) ; Rainier National Forest , Indian Flat Camp., S.E. Crumb (1, OSAC) ; Wash.Exp. Sta. Mission , 5.V.1911 (1, USNM) . Wyoming: Carbon Co., Van Dyke collection (1, CASC) . Johnson Co.: Barnum , 4.VI.1982, B.F. & J.L. Carr (1, CNCI) ; Buffalo , 12.9 km W., 20.VI.1968, S. L.W. (1, CWOB) . Lincoln Co., Frontier , 6.4 km N.W., 26.VI.1966, W. Gagne, at night (2, CWOB) . Sublette Co., Big Sandy Reservoir , 25.VI.1982, B. F. & J. L. Carr (1, CNCI) . Sweetwater Co., Green River , 1889 Wickham (1, CASC) . Teton Co., Grand Teton National Park, Gros Ventre Camp. , 24.VI.1966, W. Gagne, at night (2, CWOB) ; Grand Teton National Park, Jackson Hole Research Station , 2040 m, 4.VIII.1967, H.E. & M.A. Evans (3, USNM) ; Teton , 13.V.1931, A.R. Rolfs (1, USNM) . County unknown: “Nat.Pk., Wyo”, 1.VIII, Hubbard & Schwarz (1, USNM) ; “ Wy. ”, C. Palm collection (1, AMNH) ; Yellowstone National Park , 19.VII.1945, Bunchgrass-sage (1, USNM) ; State unknown: 22.VI.1903, P. B.G. (1, UASM) .

Figures

Plate 6. Panscopus aequalis, dorsal habitus. From Bright and Bouchard (2008), used with permission.

Map 24. Geographical distribution of Panscopus aequalis.

Figs. 17–26. Panscopus species, sternum 8 (spiculum ventrale), ventral (a) and left lateral (b) views. 17) P. torpidus; 18) P. ovalis; 19) P. tricarinatus; 20) P. bufo; 21) P. schwarzi; 22) P. coloradensis; 23) P. aequalis; 24) P. squamifrons; 25) P. abruptus; 26) P. rugicollis. Scale bar = 1 mm.

Figs. 47–52. Panscopus species, coxites and vagina, dorsal (a), ventral (b), and left lateral (c) views. 47) P. schwarzi; 48) P. coloradensis; 49) P. aequalis; 50) P. squamifrons; 51) P. abruptus; 52) P. rugicollis. cx = coxites; st = stylus. Scale bar = 1 mm.