Panscopus (Nocheles) torpidus (LeConte, 1857)

Spanton, Timothy G. & Anderson, Robert S., 2016, A Taxonomic Revision of Weevils of the Genus Panscopus Schönherr (Coleoptera: Curculionidae: Entiminae: Tropiphorini), The Coleopterists Bulletin (mo 15) 70, pp. 1-86 : 43-46

Panscopus (Nocheles) torpidus (LeConte, 1857)

( Plate 4 , Figs. 17 , 43 , 66 , Map 18 )

Hylobius torpidus LeConte 1857: 55 . LeConte 1874: 456.

Nocheles torpidus ; Horn 1876: 55.

Panscopus (Nomidus) pallidus Buchanan 1927: 31, 1936: 15 . Schenkling and Marshall 1931: 40; Buchanan 1936: 15; Blackwelder 1939: 66.

Panscopus pallidus ; Hatch 1971: 291; O’ Brien and Wibmer 1982: 63.

Panscopus torpidus ; Bright and Bouchard 2008: 266.

Panscopus (Nomidus) torpidus ; Bousquet et al. 2013: 345.

Type Material. Holotype female: “Type/8416; H. torpidus , Nocheles / torpidus /(LeC)” ( MCZC). The first label is round and faded dark blue, a color-coded label to indicate that the specimen was collected in Washington or Oregon.

Additional type material of P. pallidus : Holotype ( USNM No. 28913), Kaslo, B.C. 23 June, R. P. Currie, collector. Paratypes: Kaslo, B.C., R. P. Currie Collector, various dates PARATYPE U 913, #’s 2, 3,6–8, 12, 14–18, 20–23, and 25, ( USNM) , Kaslo, B.C., R. P. Currie, collector, 23. VI, pallidus. 5, PARATYPE USNM 28913, CNCI paratype label #4193, ( CNCI) ; Big Fork, Mont., June 1,4&15,1904 PARATYPE USNM 28913, #’s 26–32, ( USNM) ; Ainsworth, B.C. 8. VI., H. G. Dyar Collector, pallidus, 33 PARATYPE USNM 28913 ( USNM)

Notes about Synonymy. The holotype of P. torpidus is a female with five ventrites, scape not extending to the hind margin of the eye (thus, relatively short), cinereous scales giving it a grey body color, prothorax widest in front of middle, and all elytral intervals setose. It is assignable to the species that Buchanan (1927, 1936), Schenkling and Marshall (1931), Hatch (1971), and O’ Brien and Wibmer (1982) referred to as P. pallidus . The second, third, and fourth specimens in the same tray as the holotype in the LeConte collection ( MCZC) are a male and two females, also attributable to P. pallidus . The fifth specimen in the same tray is a male specimen representing the species referred to as P. torpidus by Pierce (1913), Buchanan (1927, 1936), and Hatch (1971), among others. It has a scape extending to the hind margin of the eye (thus, relatively long), and the much darker, nearly black coloration of P. torpidus of recent authors.

The specimens in the Horn collection, also in MCZC, shed no light on the identity of the species to which the name P. torpidus should apply, because the tray labeled P. torpidus includes eight specimens collectively representing four species of Panscopus and one species of Dyslobus . Two of these specimens, one female and one male, are attributable to the species referred to as P. torpidus by recent authors (not P. torpidus LeConte ). The female is nearly black in color, with only four ventrites and a long scape extending to, or nearly to, the posterior margin of the eye. This is probably the specimen to which LeConte (1874) and Horn (1876) were referring in mentioning one female having one of its abdominal segments absent. LeConte’ s (1857) description of H. torpidus is too vague to determine which species he was studying when he applied this name; however, the word “ ater ” (black) suggests he was looking at a specimen of P. torpidus of recent authors.

Horn’ s (1876) redescription of Nocheles torpidus (LeConte) is also inconclusive, but his phrase “... densely covered with cinereous scales” suggests that he was looking at material of what has recently been called P. pallidus Buchanan. Regardless , the identity of the holotype is sufficient to establish that P. pallidus Buchanan is conspecific with P. torpidus (LeConte) . This leaves without a name the species to which Pierce (1913), Buchanan (1927, 1936), Hatch (1971) and others have erroneously applied the name P. torpidus (not LeConte 1857). This species was given the name P. spantoni by Bright (Bright and Bouchard 2008).

Specific Epithet. The adjective “ torpidus ” comes from the Latin of torpid meaning numb. This probably refers to the frequent behavior of individuals of this and other species of Panscopus to remain motionless for long periods of time.

Diagnosis. The subgeneric characters listed above, plus the following distinguish specimens of this species from specimens of other species of Panscopus : straight tibia 3 straight, without dark colored spines on ventral margin; all elytral intervals setose; prothorax widest in front of middle; vestiture ashen grey in color; and rostral carinae reduced and somewhat obscured by vestiture, in many individuals appearing unicarinate.

Description. Females: Length 6.2–7.1 mm (n = 10); width across elytra 2.8–3.1 mm (n = 10). Males: Length 5.7–6.7 mm (n = 10). Color: Greybrown, vestiture of scales predominantly pale grey with patches of beige scales on dorsum, varying in size among individuals; integument, where visible, rufescent. Head: Transverse depression at base of frons present, frons and rostrum slightly discontinuous in lateral view; rostrum with indistinct medial longitudinal carina, obscured by vestiture in some specimens; nasal plate evident, without vestiture; v-shaped carina indistinct; frons and rostrum with vestiture of dense, appressed scales and scattered, decumbent, flattened setae; loose group of setae supraorbitally. Antennae: Scape relatively short, extending to anterior margin of eye; integument obscured by vestiture of dense, appressed scales and scattered, decumbent setae; funiculus (antennomeres 2–8) with vestiture of long primary setae on apical half of each article, and short, fine, appressed secondary setae on each article, integument visible. Pronotum: Approximately 1/5 wider than long in females; about 1/8 wider than long in males; median longitudinal sulcus evident, broader and deeper in anterior half; surface of pronotum rugose-tuberculate; vestiture of dense scales and decumbent setae inserted upon apices of tubercles. Elytra: Broadly oval in females; combined width about 75% of length and approximately 1/2 longer than pronotum in females; males narrower, combined width about 70% of length, on average, and 1/3 wider than pronotum; declivity, in lateral view, evenly curved to apex of elytra; alternate intervals raised above level of even intervals, more so in males; alternate intervals with partly double row of setae, even intervals with incomplete row, or nearly complete row of setae in some specimens; serial pits each with round scale, larger than surrounding scales; vestiture elsewhere of dense overlapping scales. Legs: Tibiae 1 and 2 with a few pale, stout spines on ventral margin in apical half; tibia 3 nearly straight or very slightly curved; corbellar area indistinctly closed. Abdomen: With 5 ventrites in both sexes; ventrite 1 concave at middle in males, convex at middle in females. Genitalia: Females ( Figs. 17 , 43 ): Sternum VIII with apical expansion angular in ventral view, and approximately 1/3 total length of sternum; vagina with pair of curved sclerites anteriorly; spermatheca robust, curved, and sickle-shaped, broad and thickened throughout. Males ( Fig. 66 ): Aedeagus with median lobe decurved; apex broadly rounded with weak suggestion of median apical acuity; manubrium of tegmen slightly longer than median lobe; median struts of aedeagus much longer than median lobe; internal sac membranous, finely granular in texture distad belt of tegmen; with round, horseshoe-shaped structure and a small v-shaped sclerite anteriorly in inverted sac.

Life History. Most of the approximately 1,100 adult specimens examined were collected during the period May–July. A few specimens were collected during April and August. Three specimens were collected during October, including one which was sifted from moss and one found in wood chips in March.

Three larvae (USNM) identified as P. torpidus bear labels indicating collection at Puyallup, Washington, 20 October 1934 and that they were reared. Thus, it is not certain at what stage of their development they were collected. Also, the species identification might refer to P. torpidus (LeConte) or to the one described as P. spantoni , which had erroneously been called P. torpidus by Pierce (1913) and other authors.

Plant Associations. Plant names listed on specimen labels include Pinus contorta Douglas ex Loudon , Pinus monticola Douglas ex D. Don , Pinus ponderosa P. Lawson and C. Lawson ( Pinaceae ), “ Pseudotsuga taxifolia ”, Populus tremuloides Michx. , Populus trichocarpa Torr. and A. Gray (Salicaceae) , Betula occidentalis Hook. , Acer glabrum Torr. , Salix sp. (Salicaceae) , Vaccinium sp. , “ Ribes nutkana ” ( Rubus nutkanus Moc. ex Ser. ), Spiraea discolor Pursh. (Rosaceae) , “Cherry”, Shepherdia canadensis (L.) Nutt ( Elaeagnaceae )., Ceanothus sp. (Rhamnaceae) , Symphoricarpos albus (L.) S.F. Blake ( Caprifoliaceae ), and Rosa sp. (Rosaceae)

Geographical Distribution. Weevils of this species occur in southern British Columbia, southwestern Alberta, the mountains of western Montana, northern Idaho, and much of Washington State as far west as the Cascade Mountains ( Map 18 ). Elevation records from label data vary from 610 to 1,830 m .

Chorological Affinities. The geographical ranges of P. torpidus and P. ovalis overlap one another broadly.

Material Examined. In addition to the type material listed above, more than 1,070 specimens were examined. CANADA: Alberta: Calgary, 5.VI.1959, 13.VII.1957, 28.VII.1957, 6.VI.1963, 1.VI.1972, B.F. & J.L. Carr (5, CNCI); Coleman, 28.VII.1958, 5.VII.1981, B.F. & J.L. Carr (2, CNCI); Crow’ s Nest Pass, 9.VI.1930, 17.VI.1930, J.H. Pepper (2, CNCI); Ghost Dam, 7.VII.1976, B.F. & J. L. Carr (1, CNCI); Porcupine Hills, 18.X.1959, B.F. & J.L. Carr (1, CNCI); Tp. 21 Rge. 25, W. 4 Mer., 26.V.1978, B.F. & J. L. Carr (1, CNCI); Tp. 22 Rge. 9, W. 5 Mer., 28.V.1961, B.F. & J.L. Carr (2, CNCI); Tp. 28 Rge.9, W. 5 Mer., 28.V.1961, B.F. & J.L. Carr (1, CNCI); Waterton, 7.VII.1924, H.L. Seamans (3, CNCI). British Columbia: Aspen Grove, 13.V.1934, H.B. Leech, Populus tremuloides (3, CASC; 4, SMDV); Aspen Grove, 18.V.1931, H. Richmond, Pinus ponderosa (2, CASC); Aspen Grove, 24.V.1934, N.M. Downie (1, CWOB); Aspen Grove, 1.VI.1933, K. Graham (2, SMDV); Aspen Grove, Richmond, 30.V.1931 (1, SMDV); Bx. Mt., 2.VI.1931, R. Hopping (1, SMDV; 1, CNCI); Baynes L., 3.VII.1981, 12.V.1982, B.F. & J.L. Carr (2, CNCI); Boswell, 9.V.1958, H. & A. Howden (1, CNCI); Canal Flat, 13.V.1982, B.F. & J.L. Carr (1, CNCI); Carlin, 3.VI.1978, B.F. & J.L. Carr (1, CNCI); Christina L., 6.4 km E., 2.VI.1973, G.G.E. Scudder (2, SMDV); Cranbrook, 31.V.1927, A.A. Dennys (3, CASC; 9, CNCI; 1, SMDV); Cranbrook, 13.V.1982, B.F. & J.L. Carr (2, CNCI); Creighton Valley, 10.V.1928, E.A. Rendell (1, CASC); Creston, 12.VII.1923, C.S. Lallamand (1, CASC); Creston, 27.V.1949, G. Stace-Smith, “ Rubus nutkana ” ( Ribes ) (9, CASC); Creston, 11.VI.1923, 4.VI.1923, C.B. Twigg (3, CASC); Creston, 4.VI.1923, C.B. Twigg (4, UAIC); Creston, 11.V.1958, H. & A. Howden, Populus trichocarpa (2, CNCI); Creston, 26.V.1936, G. Stace-Smith (2, CNCI); Creston, 23.V.1984, B.F. & J.L. Carr (6, CNCI); Creston,. V.1949, G. Stace-Smith (10, OSAC); Creston, 2.VI.1948, G. Stace-Smith (1, OSAC); Creston, 24.IV.1930, 5.V.1932, 7.V.1932, 1.VII.1932, 25.VI.1948, 27.VI.1948, 1.X.1953 G. Stace-Smith (11, SMDV); Creston, 4.V.1946, G. Stace-Smith, Spiraea discolor (= Holodiscus discolor ) (1, SMDV); Creston, 15.VI.1951, G. Stace-Smith, Populus trichocarpa (1, SMDV); Creston, 1.X.1953, G. Stace-Smith, sifting moss (2, SMDV); Creston, 4.V.1946, G. Stace-Smith, Acer glabrum (1, SMDV); Creston, 27.VI.1948, G. Stace-Smith, ex: Rubus nutkana (1, SMDV); Creston, 2 mi N., 12.VI.1962, C.W. O’ Brien (1, CWOB); Creston, 610 m, 20.III.1933, G. Stace-Smith, hibernating in wood chips (1, SMDV); Creston, Goat Mountain Lake, 1460 m, 1.VII.1933, G. Stace-Smith, ex: Salix (1, SMDV); Creston, Goat Mountain Lake, 1520 m, 2.VII.1933, G. Stace-Smith, Vaccinium sp. (1, SMDV); Creston, Goat Mountain Lake, 1520 m, 6.VI.1937, G. Stace-Smith (3, SMDV); Creston, Goat Mountain, Summit, 1520 m, 2.VII.1933, G. Stace-Smith (1, SMDV); Eagle River, 26.V.1932, Buckel (1, CASC); Elko, E. Kootenay, 9.VII.1949, H.B. Leech (1, CASC); Erie, 11.VI.1984, B.F. & J.L. Carr (1, CNCI); Fernie, 2.VI.1935, H.B. Leech (2, CASC); Fernie, 20.VI.1936, H.B. Leech (1, CNCI); Fernie, 24.VII.1977, B.F. & J.L. Carr (1, CNCI); Fife, 5 mi E., 8.VI.1959, R.E. Leech (1, CNCI); Fort Steele, 2.V.1977, B.F. & J.L. Carr (2, CNCI); Ft. Steele, 14.VI.1926, R. Hopping (1, CASC); Golden, 27.VI.1976, B.F. & J.L. Carr (18, CNCI); Golden, 26.IX.1976, B.F. & J.L. Carr (1, CNCI); Grand Forks, Jun-09, M. Miller (2, USNM); Gray Creek, 6.VI.1961, G. G. E. Scudder (1, SMDV); Howser, Selkirk Mountains., 22.VI.1905, J.C. Bradley (2, CUIC); Kamloops, 21.VI.1936, G.J. Spencer (1, SMDV); Kikomun Creek Provincial Park, 27.VI.1980, R.S. Anderson (2, CMNC); Kimberly, 12.VI.1926, R. Hopping (2, CASC); Larkin, 6.VII.1934, A. Thrupp (1, CASC); Lorna, 30.VI.1926, H. Richmond, Pinus contorta (1, CASC); Lumby, 28.V.1933, A. Thrupp (1, CASC; 2, CNCI); Mara, 20.VI. (1, CASC); Marysville, 3.VI.1958, 5.VI.1958, 29.VI.1958, F. Fodor (5, SMDV); Merritt, Midday Valley, 28. V.1924, Auden (1, CNCI); Nelson, 19.V.1926, R. Hopping, on Cherry (2, CASC); Nelson, 7.VI.1927, A.A. Dennys (1, CASC); New Denver, 4 mi N., 12.VIII.1966, D.S. Horning, Jr. (1, CWOB); Newgate, 2& 3. VIII.1956, J. Grant (2, SMDV); Nicola, 10.VII.1932, G.J. Spencer (1, SMDV); Oliver, Mcintyre Rd. 910 m, 1.VI & 29.V.1958, H. & A. Howden (2, CNCI); Osoyoos 1070 m, 1.VII.1953, D.F. Hardwick (1, CNCI); Paul Lake, 24.V.1933, A. Thrupp (3, CASC; 4, CNCI; 3, SMDV); Paulson, 12.VII.1962, B.F. & J. L. Carr (1, CNCI); Penticton, 4.VI.1958, H. & A. Howden (1, CNCI); Radium, 22.V.1983, B.F. & J. L. Carr (5, CNCI); Radium, 19.V.1982, B.F. & J.L. Carr (1, CNCI); Richmond, Aspen Grove, 30.V.1931 (1, CNCI); Robson, various dates-V-VII, 1947-1950, H.R. Foxlee (81, CNCI); Rock Ck., 4.8 km W., 7.VI.1959, E. E. MacDougal (1, CNCI); Rossland, 29.VII.1942, T. Laughton (1, BMNH); Salmo, 30. VI.193. VII.1980, R.S. Anderson, open riparian (1, CMNC); Salmon Arm, 2.V.1933, H.B. Leech (1, AMNH); Salmon Arm, 9.VI.1940, H.B. Leech (1, CASC); Salmon Arm, VI.1933, H.B. Leech (2, CNCI); Salmon Arm, 13.V.1932, H.B. Leech (3, SMDV; 1, UCDC; 1, USNM); Salmon Arm, 21.VI.1929, H.B. Leech (1, UCDC); Salmon Arm, 13.V.1932, H.B. Leech (1, UCDC); Salmon Arm, 13.V.1932, H.B. Leech, ex: Salix leaves (1, USNM); Sanca, 26.IV.1933, G. Stace-Smith, on Cedar wall (1, SMDV); Sanca, 26.V.1933, G. Stace-Smith, Betula occidentalis (2, SMDV); Sentinel Mountain, 1-3.VIII. 1947, H.R. Foxlee (2, CNCI); Sicamous, 2.VI.1978, B.F. & J.L. Carr (2, CNCI; 2, CMNC); Silver Star Provincial Park, 8.VIII.1973, R.H. Parry (1, CMNC); Sinclair Canyon, Kootenay National Park, 14.VI.1962, C.W. O’ Brien, Shepherdia Canadensis (1, BMNH; 4, CWOB; 1, FMNH); Six Mile Creek, 30.V.1934, A.C. Thrupp, 1 ( CASC); Stag Leap Creek 1555m elev., 12.VII.19, R.J. Cannings (1, SMDV); Tappen, 3.VI.1978, B.F. & J.L. Carr (5, CNCI); Toby Creek, 17.V.1984, B.F. & J.L. Carr (1, CNCI); Trinity Valley, 10.VI.1928, J.R. Howell, Pseudotsuga taxifolia (1, CASC); Trinity Valley, 20.V.1928, J. Howell, Pinus monticola (1, CASC); Trinity Valley, various dates, V-VI, 1928-1930, J.R.L. Howell (2, CASC; 4, CNCI; 6, SMDV); Trinity Valley, 9.V.1928, W.G. Mathers (1, USNM); Vernon, various dates V-VI, 1924-29, R. Hopping (4, CASC; 1, CMNC; 1, CNCI; 8, SMDV; 3, USNM); Vernon, 24.V.1923, E. P. Venables (1, CNCI); Vernon, 29.V.1956, R.E. Leech (1, SMDV); Vernon, 20.VI.1927, I.J. Ward (1, SMDV); Vernon, 12-May, Venables (1, USNM); Wynndel, 28.V.1950, G. Stace-Smith, Pinus contorta (2, CASC); Wynndel, 16.VI.1946, 7.VIII.1948, G. Stace-Smith (2, CASC); Wynndel, 6.VI.1948, G. Stace-Smith (1, CNCI; 1, SMDV); Wynndel, 8-15.V.1958, H. & A. Howden (12, CNCI); Wynndel, head of Lizard Creek 790 m, 4.VIII.1957, G. Stace-Smith (1, SMDV). USA: Idaho: Bonner Co., Coolin, Priest Lake, 17-23.VII.1927, E.C. Van Dyke (51, CASC); Elmira, 24.V.1984, B.F. & J.L. Carr (1, CNCI); Granite, 1.6 km W, 8.VI.1971, W.F. Barr (39, UICM); Granite, 1.6 km W, 9.VI.1972, W.F. Barr (27, UICM); Laclede, Pend Oreille River, at night, 4.VII.1966, W. Gagne, Shepherdia canadensis (2, CWOB); Little Round Lake State Park., 10. VI.1965 (1, CWOB); Little Round Lake State Park, 10.VI.1963, C.W. O’ Brien (1, CWOB); Priest Lake, 1897 (1, FMNH); Sagle, 4.VI.1950, N.M. Downie (1, FMNH); Samuels, 28.V.1932, C. Wakeland (2, USNM); Samuels, 3.2 km E., 9.VI.1972, D.E. Foster, Ceanothus (18, UICM); Samuels, Walsh, 18.VIII.1968, A.R. Gittins (2, UICM); Sandpoint, various dates, N.M. Downie (9, AMNH; 11, CNCI; 218, CWOB; 9, CMNC; 38, FMNH; 1, OSUC; 3, PURC; 1, RHTC; 3, TAMU; 3, UCDC; 9, USNM), Sandpoint, VI.1968, W.F. Barr (2, UICM); Sandpoint, 8 km E., 1.VII.1960, W.H. Lange (1, UCDC); Trout Creek, 19 km N.E. Sandpoint, 18.V.1966, W.F. Barr (2, OSAC; 1, UICM); Brush Lake, 1.VI.1969, L. S. Hawkins (1, UICM). Boundary Co., Deep Creek State Park, 12.VI.1962, C.W. O’ Brien (1, CWOB); Eastport, N.M. Downie (3, PURC). Elmore Co., Pine, 30. V.1931, H.P. Lanchester (1, AMNH). Kootenai Co., Athol, 1.VI.1967, W.F. Barr (4, UICM); Athol, 6.4 km W., 9.VI.1971, W.F. Barr, Ceanothus (3, UICM); Athol, 6 mi. W., 1.VI.1961, W.F. Barr (1, CMNC); Beauty Creek, nr. Lake Coeur D’ Alene, 4.VI.1974, W.F. Barr (4, UICM); Chilco, 2.VI.1959, W. F. Barr (2, OSAC); Farragut State Park, 23.V.1986, W. F. Barr, sweeping (1, UICM); Farragut State Park, 1.6 km S., 23.VI.1976, M. W. Hanks (6, UICM); Garfield Bay, 23.VI.1968, L.S. Hawkins, Jr. (3, UICM); Sheep Springs, 1.VI.1972, D.E. Foster (8, UICM); Thomson Creek, 4.VI.1974, W.F. Barr (1, UICM); Twin Falls (1, CMNC); Twin Lakes, 19.V.1963, G.P. Markin (1, OSAC). Shoshone Co., Clarkia, 6.4 km N.W., 23.VI.1976, D.S. Hart (2, UICM). Valley Co., McCall, Brundage Mountain, 1830 m, 24.VI.1938, M.C. Lane (1, OSAC); Snowbank Mountain, 14.VII.1976, G.A. Shook (1, CIDA). Montana: Flathead Co., Big Fork, 14.VI.1904 (3, USNM); Echo Lake, 22.V.1935, R. D. Eichman (16, MTEC); Echo Lake, N.E. Big Fork, 30.V.1935, Eichman (15, MTEC); Flathead Co.: 24.V.1936 (12, MTEC). Glacier Co., Glacier National Park, Two Medicine Lake, 1.VII.1930 (1, CASC). Lake Co., W. Flathead Lake, 15.VII.1955, R.C. Froeschner (1, MTEC); Flathead Lake, East Shore, 1.V.1934, R.D. Eichman (11, MTEC); Flathead Lake, East Shore, 1.V.1934, R.D. Eichman, on Black Raspberry (1, MTEC). Lincoln Co., Libby, 3.V III.1957, R.C. Froeschner (1, MTEC); Noxon, 16.VI.1971, N.M. Downie (1, CWOB). Ravalli Co., 19.V.1936 (1, MTEC). Sanders Co., Thomson Falls, 1 0- 23.VIII.1950, B. Malkin (2, FMNH); Unknown co.: “Koo-Koo Sint. Ridge, Cabinet National Forest, 1070-1220 m, 5.VII.1950, B. Malkin (1, FMNH). Washington: Chelan Co., Leavenworth, 13 km S.W. 8 Mile Campground., 28.VI.1987, T.G. Spanton, ex Symphoricarpos albus primarily, also Rosa , Holodiscus discolor , and Rubus sp. , at night (12, CMNC); Peshastin, 19.V.1935, W.W Baker (1, UNSM; 1, OSAC); Molson, 3.2 km N.E., 14-25.VI.1919, A.C. Burrill (1, USNM). Spokane Co., Mt. Spokane 1520– 1770 m, 22.VI.1935, K.E. Gibson (1, UICM); Mt. Spokane State Park. 1790 m, 12.VII.1971, N.M. Downie (1, CMNC). Stevens Co., Bossburg, 11.VI.1984, B.F. & J.L. Carr (1, CNCI); Lost Lake, 18.VI.1954, B. Malkin & D. Boddy (1, FMNH). Yakima Co., Mt. Rainier National Park, Sawmill Flat, 25.V.1935 (1, OSAC); Rainier National Forest, Indian Flat Camp, 5.VII.1933, S.E. Crumb (1, UNSM; 1, OSAC); Rainier National Forest, Indian Flat Camp, 27.VI.1939, S.E. Crumb (2, OSAC); Rainier National Forest, Sawmilll Flat, 15.VI.1935, J. Wilcox (4, UCDC); Sawmill Flat Forest Camp, 6.VII.1935, J. Wilcox (14, CASC); Sawmill Flat, Rainier National Forest, 25.V.1935, J. Wilcox (54, OSAC); Sawmill Flat, Rainier National Forest, 1.VI.1935, J. Wilcox (12, OSAC); Sawmill Flat, Rainier National Forest., 5.VII.1933, S.E. Crumb (2, OSAC); Sawmill Flat, Rainier National Forest, 30.V.1936, 6.VI.1937, W.W. Baker (3, OSAC); Tieton, 13.V.1931, A. R. Rolfs (1, OSAC).

Figures

Plate 4. Panscopus torpidus, dorsal habitus. From Bright and Bouchard (2008), used with permission.

Map 18. Geographical distribution of Panscopus torpidus.

Figs. 17–26. Panscopus species, sternum 8 (spiculum ventrale), ventral (a) and left lateral (b) views. 17) P. torpidus; 18) P. ovalis; 19) P. tricarinatus; 20) P. bufo; 21) P. schwarzi; 22) P. coloradensis; 23) P. aequalis; 24) P. squamifrons; 25) P. abruptus; 26) P. rugicollis. Scale bar = 1 mm.

Figs. 41–46. Panscopus species, coxites and vagina, dorsal (a), ventral (b), and left lateral (c) views. 41) P. bakeri;

Figs. 65–67. Panscopus species, aedeagus, dorsal (a), ventral (b), and left lateral (c) views. 65) P. longus; 66) P. torpidus; 67) P. ovalis. b = belt of tegmen; is = internal sac; mb = manubrium of tegmen; ml = median lobe; ms = median strut; t = tegmen. Scale bar = 1 mm.