Tridyminae Thomson, 1876

Mitroiu, Mircea-Dan, Andriescu, Ionel & Manic, Gheorghe, 2024, The Afrotropical and West-Palaearctic species of Ecrizotes Förster (Hymenoptera: Pirenidae), European Journal of Taxonomy 970, pp. 1-37 : 3

Subfamily Tridyminae Thomson, 1876

Tridymina Thomson, 1876 .

Type genus: Tridymus Ratzeburg, 1848 ; treated as Tridyminae by Ashmead (1904).

Diagnosis

Body usually with metallic reflections (with some yellowish, brown or black exceptions). Female antenna with 1–2 microscopic, 1–2 anelliform, and 3–5 large flagellomeres before a 4-segmented clava (with 3 large clavomeres and a ‘terminal button’). Eyes at most slightly divergent ventrally. Marginal vein at most 3× as long as the relatively long stigmal vein (except Ecrizotomorpha alternativa – see below, and two new species described herein where the marginal vein is slightly longer). Petiole short, hardly visible, without dorsal lamina.

Taxonomic comments

Ecrizotomorpha was previously included in the subfamily Pireninae ( Burks et al. 2022) , based on the smaller number of large flagellomeres before clava (only three instead of four or five in Tridyminae ), and the short stigmal vein found in E. alternativa . However, in other species such as E. taskhiri the fore wing venation is very similar to that of Ecrizotes and Spathopus . Apart from the peculiar structure of the flagellum, with one anelliform flagellomere flanked by two large ones ( Figs 1C , 2E , 9D , 11C , 13F ), all other characters of Ecrizotomorpha are indistinguishable from those of Ecrizotes and Spathopus . Moreover, at least in Ecrizotes monticola the same flagellomere shows various degrees of reduction. Thus, Ecrizotomorpha should be included in the subfamily Tridyminae , but the structure of the antenna alone cannot justify its distinct generic status (see below).

Figures

Fig. 1. Ecrizotes acersp. nov. A–C, E–F. Holotype, ♀ (NHMUK). D. Paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Head, frontal view. C. Antenna (arrow indicates Fu3). D. Clypeus. E. Mesosoma, dorsal view. F. Fore wing, dorsal view.

Fig. 2. Ecrizotes brevicauda Mitroiu sp. nov. A, C–E, G–H. Holotype, ♀ (NHMUK). B, F. Allotype, ♂ (NHMUK). A. Habitus, lateral view. B. Habitus, lateral view. C. Head, frontal view. D. Clypeus. E. Antennae (arrow indicates Fu3). F. Antenna. G. Mesosoma, dorsal view. H. Fore wings.

Fig. 9. Ecrizotes longus Mitroiu sp. nov.A, F–G. Holotype, ♀ (NHMUK). B, E. Allotype, ♂ (NHMUK). C–D. Paratype, ♀ (NHMUK). A. Habitus, lateral view. B. Habitus, lateral view. C. Head, frontal view. D. Antenna (arrow indicates Fu3). E. Antenna. F. Mesosoma, dorsal view. G. Fore wing, dorsal view.

Fig. 11. Ecrizotes monticola Förster, 1861. A–F. ♀ (MICO UCRC_ENT 00486453). A. Habitus, lateral view. B. Head, frontal view. C. Antenna (arrow indicates Fu3). D. Clypeus. E. Mesosoma, dorsal view. F. Fore wing, dorsal view. G–H. Female paralectotype (arrow indicates Fu3) (NMPC). I. Original label of paralectotype reading “Roseggthal” in Förster’s handwriting.

Fig. 13. Ecrizotes rovumae Mitroiu sp. nov., holotype, ♀ (MNHN). A. Habitus, lateral view. B. Head, frontal view. C. Fore wing, dorsal view. D. Clypeus. E. Mesosoma, dorsal view. F. Antenna, lateral view (arrow indicates Fu3).